Classification: Coccolithophores -> Coccolithales -> Calcidiscaceae -> Hayella
Sister taxa: Calcidiscus, Hayaster, Oolithotus, Umbilicosphaera, Cryptococcolithus, Hayella,

Distinguishing features: Coccoliths cylindrical, with shields separated by an elevated tube

Daughter taxa (blue => in age window 0-300Ma) Granddaughter taxa
Medium sized, tubular murolith, with proximal and distal flanges.

Flaring, trumpet-shaped coccolith (holotype side view only).

Medium-sized (5-7 µm) circular placoliths with an open, vacant central-area. The shield elements are distinct and show obliquity. The shield image is brighter towards the inner edge.

Medium-sized circular placolith coccoliths with a wide, open central area and highly birefringent tube cycle. Shield elements are distinct and show obliquity.

Specimens which cannot be assigned to established species


Citation: Hayella Gartner, 1969
Rank: Genus
Type species: Hayella situliformis
Synonyms: Nannocorbis Müller 1974
Notes & discussion: As explained, in detail, on the relevant Farinacci catalog page the taxon Hayella Roth is a junior homonym of Hayella Gartner.

Farinacci & Howe catalog pages: Hayella + * Nannocorbis *

Distinguishing features: Coccoliths cylindrical, with shields separated by an elevated tube

Search data:
TagsLITHS: placolith, tubular, circular, RIM: rim-high, CA: vacant,
CSPH: spherical, monomorphic, V-UNITS: distal shield, tube, R-UNITS: prox shield, DETAILS: rim-bicyclic, isogyres oblique,
MetricsLith size: 5->9µm;
The morphological data given here can be used on the advanced search page. See also these notes

Geological Range:
Last occurrence (top): within Zanclean Stage (3.60-5.33Ma, top in Zanclean stage). Data source: Total of range of species in this database
First occurrence (base): within Ypresian Stage (47.84-55.96Ma, base in Ypresian stage). Data source: Total of range of species in this database

Plot of occurrence data:


Bown, P.R.; Dunkley-Jones, T. & Young, J.R., (2007). Umbilicosphaera jordanii Bown 2005 from the Paleogene of Tanzania: confirmation of generic assignment and a Paleocene origination for the family Calcidiscaceae. Journal of Nannoplankton Research, 29(1): 25-30.

Gartner, S., (1969). Two new calcareous nannofossils from the Gulf Coast Eocene. Micropaleontology, 15: 31-34.

Gartner, S., (1969). Hayella Roth and Hayella Gartner. Micropaleontology, 15: 490.

Müller, C., (1974). Calcareous nannoplankton, Leg 25 (Western Indian Ocean). Initial Reports of the Deep Sea Drilling Project, 25: 579-633.


Hayella compiled by Jeremy R. Young, Paul R. Bown, Jacqueline A. Lees viewed: 19-3-2018

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Go to Archive.is to create a permanent copy of this page - citation notes

Comments (1)

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Rui da Gama (KL, Malaysia)

Please note: From left, 2nd and 3rd image of Hayella simplex are same as 2nd and 4th image of Hayella spp. Better to remove to avoid confusion.

Jeremy Young (UK)

Quite right - I have now changed those, but I need to check with Paul to see if all the images of Hayella sp. from Bown 2005 are actually now assignable to Hayella simplex. Thanks for pointing that out


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