Reticulofenestra pseudoumbilicus

Classification: Coccolithophores -> Isochrysidales -> Noelaerhabdaceae -> Reticulofenestra -> R. pseudoumbilicus group -> Reticulofenestra pseudoumbilicus
Sister taxa: R. calicis, R. pseudoumbilicus, R. haqii, R. minuta, R. minutula, R. asanoi, R. kahniae, R. rotaria, R. pospichalii,


Citation: Reticulofenestra pseudoumbilicus (Gartner, 1967) Gartner, 1969
Rank: Species
Basionym: Coccolithus pseudoumbilicus Gartner, 1967
Notes & discussion: For biostratigraphy it is better to separate the >7µm specimens (e.g.: Rio et al., 1990; Raffi et al., 1995); i.e. a definition of >7µm is preferable for unambiguous identification of both the NN15/16 boundary (last occurrence of R. pseudoumiblicus) and the NN10A/B boundary (start of small Reticulofenestra interval/R.pseudoumbilicus paracme). In industry it is now commonplace to use a definition of >8µm (de Kaenel pers. comm.). See Young (1990) for an extended discussion of this species.
The Palaeogene species R. dictyoda is essentially identical to R. pseudoumbilicus

Distinguishing features: Medium to large size reticulofenestrid with central opening

Farinacci & Howe catalog pages: Coccolithus pseudoumbilicus * , Reticulofenestra ampliumbilicus * , Coccolithus tenuistriatus * , Reticulofenestra pseudoumbilicus amplus *

Morphology remarks: Coccoliths >5µm (cf. e.g.: Backman, 1980; Young, 1991). Often dominates Middle Miocene - Early Pliocene assemblages.

See also: Reticulofenestra perplexa - form with closed central characteristic of the Neogene of the Southern Ocean;;

Search data:
TagsLITHS: placolith, elliptical, CA: grill, vacant,
CSPH: spherical, equant, monomorphic, V-UNITS: none, R-UNITS: all, c-str, distal shield, prox shield, tube, R-prominent, DETAILS: ca_conjunct, isogyres kinked, isogyres oblique, rim-unicyclic,
MetricsLith size: 5->12µm;
The morphological data given here can be used on the advanced search page. See also these notes

Geological Range:
Last occurrence (top): at top of NN15 zone (100% up, 3.7Ma, in Zanclean stage). Data source: zonal marker, e.g. Young 1998
First occurrence (base): within NN4 zone (14.91-17.95Ma, base in Burdigalian stage). Data source: Young 1998

Plot of occurrence data:

  • Histogram - Neptune occurrence data from DSDP and ODP proceedings. Pale shading <50 samples in time bin. Interpret with caution & read these notes
  • Note to users in China - the diagram may not plot unless you use a VPN since I use google services on the website. If this is a problem please contact me 谢谢.
  • Taxon plotted: Reticulofenestra pseudoumbilicus, synonyms included - Coccolithus pseudoumbilicus; Dictyococcites perplexus; Reticulofenestra ampliumbilicus; Reticulofenestra pseudoumbilica; Reticulofenestra pseudoumbilica (<7 microns); Reticulofenestra pseudoumbilica (>7microns); Reticulofenestra pseudoumbilica gelida; Reticulofenestra pseudoumbilica var. ampla; Reticulofenestra pseudoumbilica(5-7microns); Reticulofenestra pseudoumbilicus; Reticulofenestra pseudoumbilicus (> 7 microns); Reticulofenestra pseudoumbilica; Reticulofenestra pseudoumbilicus (>7 µm with grill);
  • space-time plot :: Samples map :: Parent: R. pseudoumbilicus group


    Bown, P. R. & Dunkley Jones, T. (2012). Calcareous nannofossils from the Paleogene equatorial Pacific (IODP Expedition 320 Sites U1331-1334). Journal of Nannoplankton Research. 32(2): 3-51. gs :: view :: open

    Bown, P. R., Gibbs, S. J., Sheward, R., O’Dea, S. & Higgins, D. (2014). Searching for cells: the potential of fossil coccospheres in coccolithophore research. Journal of Nannoplankton Research. 34(special): 5-21. gs :: view :: open

    Driever, B. W. M. (1988). Calcareous nannofossil biostratigraphy and paleoenvironmental interpretation of the Mediterranean Pliocene. Utrecht Micropaleontological Bulletin. 36: 245-. gs :: ::

    Gartner, S. (1967). Calcareous nannofossils from Neogene of Trinidad, Jamaica, and Gulf of Mexico. Paleontological Contributions, University of Kansas. 29: 1-7. gs :: ::

    Gartner, S. (1969). Correlation of Neogene planktonic foraminifera and calcareous nannofossil zones. Transactions of the Gulf Coast Association of Geological Societies. 19: 585-599. gs :: ::

    Gartner, S. (1992). Miocene nannofossil chronology in the North Atlantic, DSDP Site 608. Marine Micropaleontology. 18: 307-331. gs :: ::

    Haq, B. U. (1976). Coccoliths in cores from the Bellinghausen abyssal plain and Antarctic continental rise (DSDP Leg 35). Initial Reports of the Deep Sea Drilling Project. 35: 557-567. gs :: view :: open

    Kamptner, E. (1963). Coccolithineen-Skelettreste aus Tiefseeablagerungen des Pazifischen Ozeans. Annalen des Naturhistorischen Museums in Wien. 66: 139-204. gs :: ::

    Martini, E. (1979). Calcareous nannoplankton and silicoflagellate biostratigraphy at Reykjanes Ridge, northeastern North Atlantic (DSDP Leg 49, Sites 407 and 409). Initial Reports of the Deep Sea Drilling Project. 49: 533-549. gs :: view :: open

    Müller, C. (1974). Nannoplankton aus dem Mittel-Miozän von Walbersdorf (Burgenland). Senckenbergiana Lethaea. 55: 389-405. gs :: ::

    Müller, C. (1974). Calcareous nannoplankton, Leg 25 (Western Indian Ocean). Initial Reports of the Deep Sea Drilling Project. 25: 579-633. gs :: ::

    Theodoridis, S. (1984). Calcareous nannofossil biostratigraphy of the Miocene and revision of the helicoliths and discoasters. Utrecht Micropaleontological Bulletin. 32: 1-271. gs :: ::

    Varol, O. (1989). Calcareous nannofossil study of the central and western Solomon Islands. Circum-Pacific Council for Energy and Mineral Resources Earth Sciences. 239-268. gs :: view :: open

    Wei, W. & Wise, S. W. (1990). Middle Eocene to Pleistocene calcareous nannofossils recovered by Ocean Drilling Program leg 113 in the Weddell Sea. Proceedings of the Ocean Drilling Program, Scientific Results. 113: 639-666. gs :: view :: open

    Wise, S. W. & Wind, F. H. (1977). Mesozoic and Cenozoic calcareous nannofossils recovered by DSDP Leg 36 drilling on the Falkland Plateau, south-west Atlantic sector of the Southern Ocean. Initial Reports of the Deep Sea Drilling Project. 36(269-491): -. gs :: view :: open

    Wise, S. W. (1983). Mesozoic and Cenozoic calcareous nannofossils recovered by DSDP Leg 71 in the Falkland Plateau region, Southwest Atlantic Ocean. Initial Reports of the Deep Sea Drilling Project. 71: 481-550. gs :: view :: open

    Young, J. R. (1998). Neogene. In, Bown, P. R. (ed.) Calcareous Nannofossil Biostratigraphy. British Micropalaeontological Society Publication Series. 225-265. gs :: ::


    Reticulofenestra pseudoumbilicus compiled by Jeremy R. Young, Paul R. Bown, Jacqueline A. Lees viewed: 5-7-2020

    Taxon Search:
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    Short stable page link: Go to to create a permanent copy of this page - citation notes

    Comments (3)

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    Page 1 of 1
    Ines Galovic (Croatia)
    Isn't it Gartners' Reticulofenestra from 1969 R. pseudoumbilica, that you refer to in Citation but as pseudoumbilicus or I missed something?
    Jeremy Young (UCL, UK)
    Gartner did indeed call the taxon Reticulofenestra pseudoumbilica and this spelling was widely used. However, pseudoumibilicus is a noun not an adjective and so cannot change gender. Hence the correct spelling is pseudoumbilicus.Shirley van Heck discussed this in an article on the ICBN"""
    Ines Galovic (CGS, Croatia)
    Thank you Jeremy on explanation, I didn't check it is it a noun or not and rely on expert's knowledge, but haven't been sure till now.
    Mike Styzen (Noble Energy, US)
    The extinction or LCO of the closed central area variant/morphotype/species, usually under the name Dictyococcites antarcticus, is used in the Gulf of Mexico as a stratigraphic marker in the Early Pliocene, probably within NN 15. Wei (1990) contended that this taxon is a junior synonym of Reticulofenestra perplexa (Burns) which was described as being an order of magnitude different in size. The (informal) consensus amongst GOM workers appears to be that Burns had the unfortunate experience of describing something that did not in fact exist, and should therefore be ignored. Common usage continues to be D. antarcticus.
    Shirley van Heck (Shell International, US)
    Specimens between 5 um and 7 um have been described by Sato et al 1991 as Reticulofenestra ampla. This species has a LAD in NN16. (ref.: Sato, T., Kameo, K., Takayama, T., 1991 - Coccolith biostratigraphy of the Arabian Sea. Proc. ODP Sci. Res., 117: 37 - 54)
    Jeremy Young (NHM, UK)
    I do mention R. ampla on the minutula page, and suggest it should be regarded as a synonym of R. minutula. That is plainly not right since I also gave 5 microns as the maximum size for R. minutula. recognising these medium size reticulos in the late Pliocene is useful. There is a problem that R. ampla Sato is a junior homonym of R. ampla Theodoridis, but it would be straightforward to propose a substitute name.
    Page 1 of 1

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