Reticulofenestra umbilicus

Classification: Coccolithophores -> Isochrysidales -> Noelaerhabdaceae -> Reticulofenestra -> R. umbilicus group -> Reticulofenestra umbilicus
Sister taxa: R. umbilicus, R. dictyoda, R. hillae, R. hampdenensis, R. hesslandii, R. moorei, R. circus, R. circus var. lata, R. wadeae, R. maceria Shafik 1989, R. oamaruensis,


Citation: Reticulofenestra umbilicus (Levin, 1965) Martini & Ritzkowski, 1968
Rank: Species
Basionym: Coccolithus umbilicus Levin, 1965
Synonyms: Tremalithus placomorphus Kamptner (1948) = Reticulofenestra placomorpha (Kamptner 1948) Stradner in Haq 1968 [according to Perch-Nielsen 1985, p.543. The holotype is only 13.3 µm but the name was used for very large Reticulofenestra]

Distinguishing features: Very large (>14 µm) elliptical reticulofenestrids, with delicate net

Farinacci & Howe catalog pages: Coccolithus umbilicus *

Size: Holotype length 16.1µm.

Search data:
TagsLITHS: placolith, elliptical, CA: grill, vacant,
CSPH: spherical, equant, monomorphic, V-UNITS: none, R-UNITS: all, c-str, distal shield, prox shield, tube, R-prominent, DETAILS: ca_conjunct, isogyres kinked, isogyres oblique, rim-unicyclic,
MetricsLith size: 14->20µm;
Sources: Size range from Backman & Hermelin 1986
The morphological data given here can be used on the advanced search page. See also these notes

Geological Range:
Notes: The first appearance of very large, >14μm, elliptical reticulofenestrids (i.e. R. umbilicus) has long been used as a distinctive stratigraphic marker (e.g. Backman & Hermelin, 1986). In the Exp. 342 material these sizes are attained by subcircular to elliptical reticulofenestrids (similar to R. wadeae and R. umbilicus) in Subzone NP15b, around 3 m.y. prior to the level typically cited for this biohorizon. Agnini et al. (2014) suggest that the use of the first common occurrence of R. umbilicus may represent a more consistent horizon, above the last occurrence of C. gigas. [Bown & Newsam 2017]
Last occurrence (top): at top of NP22 zone (100% up, 32Ma, in Rupelian stage). Data source: zonal marker (see Perch-Nielsen 1985)
First occurrence (base): within NP16 zone (40.40-42.87Ma, base in Lutetian stage). Data source: Perch-Nielsen 1985

Plot of occurrence data:

  • Histogram - Neptune occurrence data from DSDP and ODP proceedings. Pale shading <50 samples in time bin. Interpret with caution & read these notes
  • Note to users in China - the diagram may not plot unless you use a VPN since I use google services on the website. If this is a problem please contact me 谢谢.
  • Taxon plotted: Reticulofenestra umbilicus, synonyms included - Apertapetra umbilica; Reticulofenestra hillae; Reticulofenestra umbilica; Reticulofenestra umbilica (>14 micron); Reticulofenestra umbilicus; Reticulofenestra umbilicus (>14um); Reticulofenestra umbilica;
  • space-time plot :: Samples map :: Parent: R. umbilicus group


    Agnini, C. et al. (2014). Biozonation and biochronology of Paleogene calcareous nannofossils from low and middle latitudes. INA Newsletter. 47: 131-181. gs

    Backman, J. & Hermelin, J. O. R. (1986). Morphometry of the Eocene nannofossil Reticulofenestra umbilicus lineage and its biochronological consequences. Palaeogeography Palaeoclimatology Palaeoecology. 57: 103-116. gs

    Bown, P. R. & Dunkley Jones, T. (2012). Calcareous nannofossils from the Paleogene equatorial Pacific (IODP Expedition 320 Sites U1331-1334). Journal of Nannoplankton Research. 32(2): 3-51. gs

    Bown, P. R. & Newsam, C. (2017). Calcareous nannofossils from the Eocene North Atlantic Ocean (IODP Expedition 342 Sites U1403–1411). Journal of Nannoplankton Research. 37(1): 25-60. gs

    Bown, P. R. (2005d). Palaeogene calcareous nannofossils from the Kilwa and Lindi areas of coastal Tanzania (Tanzania Drilling Project 2003-4). Journal of Nannoplankton Research. 27(1): 21-95. gs

    Bown, P. R., Gibbs, S. J., Sheward, R., O’Dea, S. & Higgins, D. (2014). Searching for cells: the potential of fossil coccospheres in coccolithophore research. Journal of Nannoplankton Research. 34(special): 5-21. gs

    Bybell, L. (1975). Middle Eocene calcareous nannofossils at Little Stave Creek, Alabama. Tulane Studies in Geology and Paleontology. 11(4): 177-247. gs

    de Kaenel, E. & Villa, G. (1996). Oligocene-Miocene calcareous nannofossil biostratigraphy and paleoeecology from the Iberian Abyssal Plain. Proceedings of the Ocean Drilling Program. Scientific Results. 149: 79-145. gs

    Dunkley Jones, T., Bown, P. R. & Pearson, P. (2009). Exceptionally well preserved upper Eocene to lower Oligocene calcareous nannofossils (Prymnesiophycidae) from the Pande Formation (Kilwa Group), Tanzania. Journal of Systematic Palaeontology. 7(4): 359-411. gs

    Fioroni, C., Villa, G., Persico, D., Wise, S. W. & Pea, L. (2012). Revised middle Eocene-upper Oligocene calcareous nannofossil biozonation for the Southern Ocean. Revue de Micropaléontologie. 55(2): 53-70. gs

    Fioroni, C., Villa, G., Persico, D. & Jovane, L. (2015a). Middle Eocene-Lower Oligocene calcareous nannofossil biostratigraphy and paleoceanographic implications from Site 711 (equatorial Indian Ocean). Marine Micropaleontology. 118: 50-62. gs

    Haq, B. U. (1968). Studies on upper Eocene calcareous nannoplankton from NW Germany. Stockholm Contributions in Geology. 18: 13-74. gs

    Kamptner, E. (1948). Coccolithen aus dem Torton des Inneralpinen Wiener Beckens. Anzeiger. Österreichische Akademie der Wissenschaften. Mathematische-Naturwissenschafliche Klasse. Wien. 157: 1-16. gs

    Levin, H. L. (1965). Coccolithophoridae and related microfossils from the Yazoo Formation (Eocene) of Mississippi. Journal of Paleontology. 39(2): 265-272. gs

    Martini, E. & Ritzkowski, S. (1968). Was ist das 'Unter-Oligozän'? Eine Analyse der Beyrich'schen und v Koenen'schen Fassung der Stufe mit Hilfe des fossilen Nannoplanktons. Nachrichten der Akademie der Wissenschaften in Gottingen. II. Mathematisch-Physikalische Klasse. 1968, 13: 231-250. gs

    Perch-Nielsen, K. (1971b). Elektronenmikroskopische untersuchungen an Coccolithen und verwandten Formen aus dem Eozan von Danemark. Biologiske Skrifter. Kongelige Danske Videnskabernes Selskab. 18(3): 1-76. gs

    Perch-Nielsen, K. (1972). Remarks on Late Cretaceous to Pleistocene coccoliths from the North Atlantic. Initial Reports of the Deep Sea Drilling Project. 12: 1003-1069. gs

    Perch-Nielsen, K. (1985a). Cenozoic calcareous nannofossils. In, Bolli, H. M. , Saunders, J. B. & Perch-Nielsen, K. (eds) Plankton Stratigraphy. Cambridge University Press, Cambridge 427-555. gs

    Self-Trail, J. (2011). Paleogene Calcareous Nannofossils of the South Dover Bridge core, Southern Maryland (USA). Journal of Nannoplankton Research. 32(1): 1-28. gs

    Shepherd, C. L. & Kulhanek, D. K. (2016). Eocene nannofossil biostratigraphy of the mid-Waipara River section, Canterbury Basin, New Zealand. Journal of Nannoplankton Research. 26(1): 33-59. gs

    Varol, O. (1998). Palaeogene. In, Bown, P. R. (ed.) Calcareous Nannofossil Biostratigraphy. British Micropalaeontological Society Publication Series. 200-224. gs

    Wei, W. & Wise, S. W. (1990). Middle Eocene to Pleistocene calcareous nannofossils recovered by Ocean Drilling Program leg 113 in the Weddell Sea. Proceedings of the Ocean Drilling Program. Scientific Results. 113: 639-666. gs

    Wise, S. W. (1983). Mesozoic and Cenozoic calcareous nannofossils recovered by DSDP Leg 71 in the Falkland Plateau region, Southwest Atlantic Ocean. Initial Reports of the Deep Sea Drilling Project. 71: 481-550. gs


    Reticulofenestra umbilicus compiled by Jeremy R. Young, Paul R. Bown, Jacqueline A. Lees viewed: 26-2-2020

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    Comments (1)

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    Page 1 of 1
    Simun Ascic (Zagreb University, Croatia)
    Description says <14µm. probably should be >14 µm
    Paul Bown (Univeristy College London, UK)
    Well spotted! Just fixed it.
    Page 1 of 1

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