Catapsydrax howei

Classification: pf_cenozoic -> Globigerinidae -> Catapsydrax -> Catapsydrax howei
Sister taxa: C. indianus, C. dissimilis, C. globiformis, C. africanus, C. howei, C. unicavus, C. sp.,


Citation: Catapsydrax howei (Blow and Banner 1962)
Rank: Species
Basionym: Globigerinita howei
Taxonomic discussion: The loosely coiled test with large circular, infralaminal apertures of Catapsydrax howei somewhat resembles Recent Globigerinoides, suggesting a shallow habitat in the water column. We illustrate the holotype in SEM for the first time (Pl.5.2, Figs. 9-12) along with newly collected specimens from the type locality (Pl.5.2, Figs. 13-16). Catapsydrax golicynensis Menkes from the upper Eocene of the Black Sea exhibits the large arched aperture of this species and is a probable synonym. The specimen figured by Warraich and Ogasawara (2001) is a five-chambered form that does not match the morphology of C. howei. [Olsson et al. 2006]

Catalog entries: Globigerinita howei, Catapsydrax golicynensis

Type images:

Distinguishing features: Larger, loosely coiled test, with large globular bulla with 3 large, circular, infralaminal apertures

NB These concise distinguishing features statements are used in the tables of daughter-taxa to act as quick summaries of the differences between e.g. species of one genus.
They are being edited as the site is developed and comments on them are especially welcome.


Diagnostic characters: Catapsydrax howei is distinguished from C. africanus by its larger, more loosely coiled test, large globular bulla with large, circular, infralaminal apertures. Catapsydrax globiformis is smaller, has a spherical-shaped test with a low slightly inflated bulla with small circular, infralaminal apertures. [Olsson et al. 2006]

Wall type: Cancellate, probably spinose in life, ruber/sacculifer-type wall texture. [Olsson et al. 2006]

Test morphology: Low trochospiral, lobulate test with generally 3 chambers, somewhat loosely coiled, in the final whorl. Chambers inflated, globular increasing rapidly in size with a large globular, inflated bulla covering the umbilicus; in spiral view 3 globular chambers increasing rapidly in size, sutures straight to slightly curved, moderated depressed; in umbilical view dominated by a large inflated bulla with 3 large circular, infralaminal apertures with a continuous, thickened imperforate rim facing each suture; sutures straight, somewhat strongly depressed; in edge view globular in shape with bulla showing one infralaminal aperture opening onto the suture line. [Olsson et al. 2006]

Size: Holotype maximum diameter 0.54 mm, thickness 0.53 mm. [Olsson et al. 2006]

Character matrix

test outline:Lobatechamber arrangement:Trochospiraledge view:Equally biconvexaperture:Umbilical
sp chamber shape:Globularcoiling axis:Lowperiphery:N/Aaperture border:Bulla
umb chbr shape:Globularumbilicus:Wideperiph margin shape:Broadly roundedaccessory apertures:Infralaminal
spiral sutures:Strongly depressedumb depth:Deepwall texture:Cancellateshell porosity:Finely Perforate: 1-2.5µm
umbilical or test sutures:Strongly depressedfinal-whorl chambers:3.0-3.0 N.B. These characters are used for advanced search. N/A - not applicable

Biogeography and Palaeobiology

Geographic distribution: Known from tropical to mid latitude sites. [Olsson et al. 2006]
Aze et al. 2011 summary: Low to middle latitudes; based on Olsson et al. (2006c)

Isotope paleobiology: No data available. [Olsson et al. 2006]
Aze et al. 2011 ecogroup 4 - Open ocean sub-thermocline. Based on very light δ13C and very heavy δ18O. Sources cited by Aze et al. 2011 (appendix S3): Keller (1985)

Phylogenetic relations: Catapsydrax howei probably evolved from C. unicavus in the middle Eocene by becoming larger and more loosely coiled, and developing a large, globular, inflated bulla with large circular, infralaminal apertures. [Olsson et al. 2006]

Most likely ancestor: Catapsydrax unicavus - at confidence level 4 (out of 5). Data source: Olsson et al. 2006 f5.1.
Likely descendants: Catapsydrax africanus;

Biostratigraphic distribution

Geological Range:
Notes: Zone E9 to E16. [Olsson et al. 2006]
Last occurrence (top): within E16 zone (33.90-34.68Ma, top in Priabonian stage). Data source: Olsson et al. 2006 f5.1
First occurrence (base): within E9 zone (43.23-43.85Ma, base in Lutetian stage). Data source: Olsson et al. 2006 f5.1

Plot of occurrence data:

Primary source for this page: Olsson et al. 2006 - Eocene Atlas, chap. 5, p. 72


Blow, W. H. & Banner, F. T. (1962). The mid-Tertiary (Upper Eocene to Aquitanian) Globigerinaceae. In, Eames, F. E. , Banner, F. T. , Blow, W. H. & Clarke, W. J. (eds) Fundamentals of mid-Tertiary Stratigraphical Correlation. Cambridge University Press, Cambridge 61-151. gs

Menkes, M. A. (1975). Representatives of the genus Catapsydrax in the Eocene deposits of the South Ukraine. Akad Nauk SSSR, Sib Otd, Inst Geol Geofiz, Trudy. 333: 281-. gs

Olsson, R. K., Pearson, P. N. & Huber, B. T. (2006c). Taxonomy, biostratigraphy, and phylogeny of Eocene Catapsydrax, Globorotaloides, Guembelitrioides, Paragloborotalia, Parasubbotina, and Pseudoglobigerinella n. gen. In, Pearson, P. N. , Olsson, R. K. , Hemleben, C. , Huber, B. T. & Berggren, W. A. (eds) Atlas of Eocene Planktonic Foraminifera. Cushman Foundation for Foraminiferal Research, Special Publication. 41(Chap 5): 67-110. gs

Warraich, M. Y. & Ogasawara, K. (2001). Tethyan Paleocene-Eocene planktic foraminifera from the Rakhi Nala and Zinda Pir land sections of the Sulaiman Range, Pakistan. Science Reports of the Institute of Geosciences, University of Tsukuba, Section B = Geological Sciences. 22: 1-59. gs


Catapsydrax howei compiled by the pforams@mikrotax project team viewed: 16-9-2019

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