Morozovelloides bandyi

Classification: pf_cenozoic -> Truncorotaloididae -> Morozovelloides -> Morozovelloides bandyi
Sister taxa: M. lehneri, M. coronatus, M. crassatus, M. bandyi, M. sp.,


Citation: Morozovelloides bandyi (Fleisher 1974)
Rank: Species
Basionym: Morozovella bandyi Fleisher 1974
Taxonomic discussion: Morozovelloides bandyi is a common form in Zones E8 and E9, but first appears in the upper part of Zone E7 where it intergrades with Acarinina praetopilensis. It has probably been described previously on many occasions as crassata or spinulosa, but such synonymies in the older literature are difficult to verify because M. bandyi is strongly homeomorphic to the Morozovella aequaM. subbotinae group of the lower Eocene. [Pearson & Berggren 2006]

Catalog entries: Morozovella bandyi,

Type images:

Distinguishing features: Distinguished from all other species of Morozovelloides by the strongly disjunct long axes of the chambers, as seen in spiral view.

NB These concise distinguishing features statements are used in the tables of daughter-taxa to act as quick summaries of the differences between e.g. species of one genus.
They are being edited as the site is developed and comments on them are especially welcome.


Diagnostic characters: Distinguished from all other species of Morozovelloides by the strongly disjunct long axes of the chambers, as seen in spiral view, and from Acarinina praetopilensis by having a more dorso-ventrally flattened test and peripheral (as opposed to circum-cameral) muricocarina. [Pearson & Berggren 2006]

Wall type: Muricate, normal perforate with smooth areas on the spiral side. [Pearson & Berggren 2006]

Test morphology: Planoconvex to lenticular test, coiled in a low trochospiral, with concentration of bladed muricae around the petaloid periphery that constitute a discontinuous muricocarina. Four to five chambers in the final whorl, added in such a way that their long axes are strongly disjunct when viewed on the dorsal side. Typically the chambers are imbricate, sometimes strongly so, as if tilted relative to the dorso-ventral plane. Small supplementary apertures are commonly present at the base of the sutures between the penultimate and antepenultimate chambers, and more rarely earlier in the spiral. Primary aperture is a low, flat arch, in an umbilical-extraumbilical position. Umbilicus moderately broad and deep. Ventral sutures depressed with umbilical shoulders of chambers commonly showing concentrations of muricae (but not to the extent seen in M. coronata, see below). [Pearson & Berggren 2006]

Size: Maximum diameter of holotype about 0.28 mm; height 0.16 mm. [Pearson & Berggren 2006]

Character matrix

test outline:Lobatechamber arrangement:Trochospiraledge view:Planoconvexaperture:Umbilical-extraumbilical
sp chamber shape:Crescenticcoiling axis:Lowperiphery:Muricocarinateaperture border:N/A
umb chbr shape:Subtriangularumbilicus:Wideperiph margin shape:Subangularaccessory apertures:None
spiral sutures:Raised muricateumb depth:Deepwall texture:Moderately muricateshell porosity:Finely Perforate: 1-2.5µm
umbilical or test sutures:Moderately depressedfinal-whorl chambers:4.0-5.0 N.B. These characters are used for advanced search. N/A - not applicable

Biogeography and Palaeobiology

Geographic distribution: Cosmopolitan. [Pearson & Berggren 2006]
Aze et al. 2011 summary: Cosmopolitan; based on Pearson & Berggren (2006)

Isotope paleobiology: Oxygen and carbon isotopes indicate a mixed layer, phososymbiotic habitat (Boersma and others, 1987 [recorded as Morozovella cf. aequa], Pearson and others, 1993 and Pearson and others, 2001 [recorded as Morozovella crassata]). Boron isotope data suggest a deeper habitat than co-occurring M. crassatus (Pearson and Palmer, 1999, recorded as Morozovella crassata; Morozovelloides crassatus recorded as Morozovella spinulosa). [Pearson & Berggren 2006]
Aze et al. 2011 ecogroup 1 - Open ocean mixed-layer tropical/subtropical, with symbionts. Based on very heavy δ13C and relatively light δ18O. Sources cited by Aze et al. 2011 (appendix S3): Boersma et al. (1987); Pearson et al. (1993, 2001a)

Phylogenetic relations: Morozovelloides bandyi evolved from Acarinina praetopilensis and gave rise to M. crassatus. [Pearson & Berggren 2006]

Most likely ancestor: Acarinina praetopilensis - at confidence level 4 (out of 5). Data source: Pearson & Berggren 2006, f10.1.
Likely descendants: Morozovelloides crassatus;

Biostratigraphic distribution

Geological Range:
Notes: First appears in the upper part of Zone E7, intergrading with A. praetopilensis. Most common in Zones E8 and E9, but ranging as high as Zone E10. [Pearson & Berggren 2006] NB Given the problems with zones E7 and E8 disccussed by Wade et al. 2100 this range may need revising [JRY 2017]
Last occurrence (top): within E10 zone (41.89-43.23Ma, top in Lutetian stage). Data source: Pearson & Berggren 2006, f10.1
First occurrence (base): within E7b subzone (45.72-48.31Ma, base in Ypresian stage). Data source: Pearson & Berggren 2006, f10.1

Plot of occurrence data:

Primary source for this page: Pearson & Berggren 2006 - Eocene Atlas, chap. 10, p. 330


Beckmann, J. P. (1953). Die Foraminiferen der Oceanic Formation (Eocaen-Oligocaen) von Barbados, Kl. Antillen. Eclogae Geologicae Helvetiae. 46: 301-412. gs

Boersma, A., Premoli Silva, I. & Shackleton, N. J. (1987). Atlantic Eocene planktonic foraminiferal paleohydrographic indicators and stable isotope paleoceanography. Paleoceanography. 2: 287-331. gs

Cushman, J. A. & Barksdale, J. D. (1930). Eocene foraminifera from Martinez, California. Contributions Department of Geology, Stanford University. 1: 55-73. gs

Cushman, J. A. (1925b). An Eocene fauna from the Moctezuma River, Mexico. Bulletin of the American Association of Petroleum Geologists. 9(2): 298-301. gs

Cushman, J. A. (1927b). New and interesting foraminifera from Mexico and Texas. Contributions from the Cushman Laboratory for Foraminiferal Research. 3: 111-119. gs

Cushman, J. A. (1939b). Eocene foraminifera from submarine cores off the eastern coast of North America. Contributions from the Cushman Laboratory for Foraminiferal Research. 15: 49-76. gs

Fleisher, R. L. (1974a). Cenozoic planktonic foraminifera and biostratigraphy, Arabian Sea, Deep Sea Drilling Project, Leg 23A. Initial Reports of the Deep Sea Drilling Project. 23: 1001-1072. gs

Pearson, P. N. & Berggren, W. A. (2006). Taxonomy, biostratigraphy, and phylogeny of Morozovelloides n. gen. In, Pearson, P. N. , Olsson, R. K. , Hemleben, C. , Huber, B. T. & Berggren, W. A. (eds) Atlas of Eocene Planktonic Foraminifera. Cushman Foundation for Foraminiferal Research, Special Publication. 41: 343-376. gs

Pearson, P. N. & Palmer, M. R. (1999). Middle Eocene seawater pH and atmospheric carbon dioxide concentrations. Science. 284: 1824-1826. gs

Pearson, P. N., Shackleton, N. J. & Hall, M. A. (1993). Stable isotope paleoecology of middle Eocene planktonic foraminifera and multi-species isotope stratigraphy, DSDP Site 523, South Atlantic. Journal of Foraminiferal Research. 23: 123-140. gs


Morozovelloides bandyi compiled by the pforams@mikrotax project team viewed: 20-7-2019

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