Nannotax3 - ntax_cenozoic - Toweius pertusus Nannotax3 - ntax_cenozoic - Toweius pertusus

Toweius pertusus


Classification: ntax_cenozoic -> Isochrysidales -> Prinsiaceae -> Toweius -> Toweius pertusus
Sister taxa: T. oztunalii ⟩⟨ T. occultatus, T. eminens, T. tovae, T. serotinus, T. pertusus ⟩⟨ T. reticulum, T. selandianus, T. rotundus, T. patellus, T. callosus, T. magnicrassus, T. brusselensis ⟩⟨ T. gammation ⟩⟨ T. sp.

Taxonomy:

Citation: Toweius pertusus (Sullivan, 1965) Romein, 1979
taxonomic rank: Species
Basionym: Coccolithus pertusus Sullivan, 1965
Synonyms:

Distinguishing features:
Parent taxon (Toweius): Typically medium-sized.(5-8 µm), elliptical to circular, central areas typically with a conjunct net and/or bars.
This taxon: Small to medium-sized (4-6 µm) with >=10 pores

Farinacci & Howe catalog pages: C. pertusus * , T. craticulus *


Morphology:

Small to medium-sized; elliptical to sub-circular; central area with perforate net; perforations typically small and may be difficult to resolve in LM.


Phylogenetic relations

This is the main mid Palaeocene Toweius species and gives rise to larger species in the late Palaeocene and Early Eocene

Most likely ancestor: Toweius selandianus - at confidence level 3 (out of 5). Data source: Bown et al. 2023, fig.8.
Likely descendants: Toweius tovae; plot with descendants

Search data:
LITHS: placolith, circular, elliptical, CA: ca_conjunct, grill, pores,
CSPH: equant, monomorphic, CROSS-POLARS: rim-bicyclic, V-prominent, R-prominent,
Lith size: 3->8µm;
The morphological data given here can be used on the advanced search page. See also these notes

Geological Range:
Notes: Monechi et al. (2013) reccomend base of T. pertusus as alternative marker for NP3/4 boundary
Last occurrence (top): within NP14 zone (46.29-49.11Ma, top in Lutetian stage). Data source: [PRB rough estimate]
First occurrence (base): at base of NP4 zone (0% up, 63.3Ma, in Danian stage). Data source: Monechi et al. 2013, Fig 5

Plot of occurrence data:

References:

Agnini, C., Fornaciari, E. & Raffi, I. (2008). Three new species of calcareous nannofossil from Late Palaeocene and Early Eocene assemblages (Ocean Drilling Program Site 1262, Walvis Ridge, SE Atlantic Ocean). Journal of Nannoplankton Research. 30(1): 51-56. gs

Bown, P. R. & Dunkley Jones, T. (2012). Calcareous nannofossils from the Paleogene equatorial Pacific (IODP Expedition 320 Sites U1331-1334). Journal of Nannoplankton Research. 32(2): 3-51. gs O

Bown, P. R. & Newsam, C. (2017). Calcareous nannofossils from the Eocene North Atlantic Ocean (IODP Expedition 342 Sites U1403–1411). Journal of Nannoplankton Research. 37(1): 25-60. gs O

Bown, P. R. (2005d). Palaeogene calcareous nannofossils from the Kilwa and Lindi areas of coastal Tanzania (Tanzania Drilling Project 2003-4). Journal of Nannoplankton Research. 27(1): 21-95. gs O

Bown, P. R. (2010). Calcareous nannofossils from the Paleocene/Eocene Thermal Maximum interval of southern Tanzania (TDP Site 14). Journal of Nannoplankton Research. 31(1): 11-38. gs O

Bown, P. R. (2016). Paleocene calcareous nannofossils from Tanzania (TDP sites 19, 27 and 38). Journal of Nannoplankton Research. 36(1): 1-32. gs O

Bown, P. R., Gibbs, S. J., Sheward, R., O’Dea, S. & Higgins, D. (2014). Searching for cells: the potential of fossil coccospheres in coccolithophore research. Journal of Nannoplankton Research. 34(S): 5-21. gs

Bown, P. R., Kim, H. & Gibbs, S. J. (2023). Danian calcareous nannofossil evolution and taxonomy with focus on sites from the North Atlantic Ocean (IODP Expedition 342, Sites U1403 and U1407). Journal of Nannoplankton Research. 41(2): 110-157. gs O

Bybell, L. M. & Self-Trail, J. (1995). Evolutionary, biostratigraphic and taxonomic study of calcareous nanofossils from a continuous Palaeocene-Eocene boundary section in New Jersey. U.S. Geological Survey, Professional Paper. 1554: 1-36. gs O

Hay, W. W. & Mohler, H. P. (1967). Calcareous nannoplankton from Early Tertiary rocks at Point Labau, France and Paleocene-Early Eocene correlations. Journal of Paleontology. 41(6): 1505-1541. gs

MacKenzie, D. T. & Wise,S. W. (1983). Paleocene and Eocene calcareous nannofossils from Deep Sea Drilling Project Legs 25 and 40, South and East of Africa. Initial Reports of the Deep Sea Drilling Project. 71(2): 1141-1170. gs

Mai, H., Hillebrandt-Habel, T., von Salis Perch-Nielsen, K. & Willems, H. (1998). Paleocene coccospheres from DSDP Leg 39, Site 356, Sao Paulo Plateau, S. Atlantic Ocean. Journal of Nannoplankton Research. 20(1): 21-29. gs

Monechi, S., Reale, V., Bernaola, G. & Balestra, B. (2013). The Danian/Selandian boundary at Site 1262 (South Atlantic) and in the Tethyan region: Biomagnetostratigraphy, evolutionary trends in fasciculiths and environmental effects of the Latest Danian Event. Marine Micropaleontology. 98(1): 28-40. gs

Müller, C. (1979). Calcareous nannofssils from the North Atlantic (Leg 48). Initial Reports of the Deep Sea Drilling Project. 48: 589-639. gs

Okada, H. & Thierstein, H. R. (1979). Calcareous nannoplankton - Leg 43, Deep Sea Drilling Project. Initial Reports of the Deep Sea Drilling Project. 43: 507-573. gs O

Perch-Nielsen, K. (1971b). Einige neue Coccolithen aus dem Paläozän der Bucht von Biskaya und dem Eozän der Labrador See. Bulletin of the Geological Society of Denmark. 21: 347-361. gs

Perch-Nielsen, K. (1971c). Elektronenmikroskopische untersuchungen an Coccolithen und verwandten Formen aus dem Eozan von Danemark. Biologiske Skrifter, Kongelige Danske Videnskabernes Selskab. 18(3): 1-76. gs O

Perch-Nielsen, K. (1977). Albian to Pleistocene calcareous nannofossils from the Western South Atlantic, DSDP Leg 39. Initial Reports of the Deep Sea Drilling Project. 39: 699-823. gs O

Perch-Nielsen, K., Sadek, A., Barakat, M. G. & Teleb, F. (1978). Late Cretaceous and Early Tertiary Calcareous nannofossil and Planktonic foraminifera zones from Egypt. Actes du VI Colloque Africain de Micropaléontologie - Tunis 1974. Annales des Mines et de la Géologie Tunissienne. 28: 337-403. gs

Pospichal, J. J. & Wise, S. W. (1990). Paleocene to Middle Eocene calcareous nannofossils of ODP sites 689 and 690, Maud Rise, Weddell Sea. Proceedings of the Ocean Drilling Program, Scientific Results. 113: 613-638. gs

Romein, A. J. T. (1979). Lineages in Early Paleogene calcareous nannoplankton. Utrecht Micropaleontological Bulletin. 22: 1-231. gs O

Self-Trail, J. (2011). Paleogene Calcareous Nannofossils of the South Dover Bridge core, Southern Maryland (USA). Journal of Nannoplankton Research. 32(1): 1-28. gs

Self-Trail, J. M., Powars, D. S., Watkins, D. K. & Wandless, G. A. (2012). Calcareous nannofossil assemblage changes across the Paleocene–Eocene Thermal Maximum: Evidence from a shelf setting. Marine Micropaleontology. 92–93: 61-80. gs

Shepherd, C. L. & Kulhanek, D. K. (2016). Eocene nannofossil biostratigraphy of the mid-Waipara River section, Canterbury Basin, New Zealand. Journal of Nannoplankton Research. 36(1): 33-59. gs O

Sullivan, F. R. (1965). Lower Tertiary nannoplankton from the California Coast Ranges. II. Eocene. University of California Publications in Geological Sciences. 53: 1-74. gs O

Thomsen, E. & Heilmann-Clausen, C. (1984). The Danian-Selandian boundary at Svejstrup with remarks on the biostratigraphy of the boundary in Western Denmark. Bulletin of the Geological Society of Denmark. 33: 341-362. gs O

Varol, O. (1989b). Eocene calcareous nannofossils from Sile (northwest Turkey). Revista Española de Micropaleontología. 21: 273-320. gs

Varol, O. (1998). Palaeogene. In, Bown, P. R. (ed.) Calcareous Nannofossil Biostratigraphy. British Micropalaeontological Society Publication Series . 200-224. gs

Wise, S. W. & Wind, F. H. (1977). Mesozoic and Cenozoic calcareous nannofossils recovered by DSDP Leg 36 drilling on the Falkland Plateau, south-west Atlantic sector of the Southern Ocean. Initial Reports of the Deep Sea Drilling Project. 36(269-491): -. gs O


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Toweius pertusus compiled by Jeremy R. Young, Paul R. Bown, Jacqueline A. Lees viewed: 11-10-2024

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Comments (2)

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Dave Bord(US)

The note under the Geological Range recommends the base of T. pertusus as an alternative marker for the NP2/NP3 boundary, but it should say the NP3/NP4 boundary as suggested by Monechi et al. (2013, p 31).

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Jeremy Young(UK)

Thanks David, I will change that now.

cheers Jeremy