Taxonomy:
NB The epithet protoannulus is based on the Latin masculine noun annulus (ring). It thus does not change gender and the correct orthography is Umbilicosphaera protoannulus.
Distinguishing features:
Parent taxon (Umbilicosphaera): Central area open
This taxon: Small to medium sized (4.5-8.0 µm) ring-shaped coccoliths that are distinctly bicyclic in XPL: the bright inner cycle is narrower than the outer cycle, and crossed by extinction lines that are near axial.
Farinacci & Howe catalog pages: C. protoannula * , C. pakistanica *
Search data:
LITHS: placolith, circular, elliptical, CA: vacant, CSPH: equant, monomorphic, CROSS-POLARS: rim-bicyclic, V-prominent, R-prominent, |
Lith size: 4.5->8µm; |
Geological Range:
Last occurrence (top): within NP21 zone (32.92-34.44Ma, top in Rupelian stage). Data source: Bown et al., 2007
First occurrence (base): within NP11 zone (53.70-54.17Ma, base in Ypresian stage). Data source: Bown et al., 2007
Plot of occurrence data:
Bown, P. R. & Dunkley Jones, T. (2012). Calcareous nannofossils from the Paleogene equatorial Pacific (IODP Expedition 320 Sites U1331-1334). Journal of Nannoplankton Research. 32(2): 3-51. gs O Bown, P. R. & Newsam, C. (2017). Calcareous nannofossils from the Eocene North Atlantic Ocean (IODP Expedition 342 Sites U1403–1411). Journal of Nannoplankton Research. 37(1): 25-60. gs O Bown, P. R. (2005d). Palaeogene calcareous nannofossils from the Kilwa and Lindi areas of coastal Tanzania (Tanzania Drilling Project 2003-4). Journal of Nannoplankton Research. 27(1): 21-95. gs O Dunkley Jones, T., Bown, P. R. & Pearson, P. (2009). Exceptionally well preserved upper Eocene to lower Oligocene calcareous nannofossils (Prymnesiophycidae) from the Pande Formation (Kilwa Group), Tanzania. Journal of Systematic Palaeontology. 7(4): 359-411. gs Gartner, S. (1971). Calcareous nannofossils from the JOIDES Blake Plateau cores and revision of the Paleogene nannofossil zonation. Tulane Studies in Geology and Paleontology. 8: 101-121. gs O Haq, B. U. & Lohmann, G. P. (1976). Early Cenozoic calcareous nannoplankton biogeography of the Atlantic Ocean. Marine Micropaleontology. 1: 119-194. gs Jones, A. P. & Dunkley Jones, T. (2020). Middle Eocene to Early Oligocene calcareous nannofossils from the Nanggulan Formation, Java, Indonesia. Journal of Nannoplankton Research. 38(1): 57-79. gs Loeblich, A. R. & Tappan, H. (1978). The coccolithophorid genus Calcidiscus Kamptner and its synonyms. Journal of Paleontology. 52: 1390-1392. gs Müller, C. (1979). Calcareous nannofssils from the North Atlantic (Leg 48). Initial Reports of the Deep Sea Drilling Project. 48: 589-639. gs Perch-Nielsen, K. (1977). Albian to Pleistocene calcareous nannofossils from the Western South Atlantic, DSDP Leg 39. Initial Reports of the Deep Sea Drilling Project. 39: 699-823. gs O Shamrock, J. L. & Watkins, D. K. (2012). Eocene calcareous nannofossil biostratigraphy and community structure from Exmouth Plateau, Eastern Indian Ocean (ODP Site 762). Stratigraphy. 9(1): 1-11. gs Shepherd, C. L. & Kulhanek, D. K. (2016). Eocene nannofossil biostratigraphy of the mid-Waipara River section, Canterbury Basin, New Zealand. Journal of Nannoplankton Research. 36(1): 33-59. gs O Young, J. R. & Bown, P. R. (2014). Some emendments to calcareous nannoplankton taxonomy. Journal of Nannoplankton Research. 33(1): 39-46. gs References:
Umbilicosphaera protoannulus compiled by Jeremy R. Young, Paul R. Bown, Jacqueline A. Lees viewed: 16-9-2024
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