Toweius occultatus


Classification: ntax_cenozoic -> Isochrysidales -> Prinsiaceae -> Toweius -> Toweius occultatus
Sister taxa: T. serotinus, T. pertusus, T. tovae, T. eminens, T. occultatus ⟩⟨ T. reticulum, T. selandianus, T. rotundus, T. patellus, T. callosus, T. magnicrassus ⟩⟨ T. gammation ⟩⟨ T. sp.

Taxonomy:

Citation: Toweius occultatus (Locker, 1967) Perch-Nielsen, 1971
Rank: Species
Basionym: Coccolithus occultatus Locker, 1967
Synonyms: Coccolithus petrinus Stradner, 1969 [my identification - JRY 2015]

Distinguishing features: Medium to large with 2 large openings in the central area.

Farinacci & Howe catalog pages: C. occultatus * , C. petrinus *


Morphology: Medium to large; elliptical to sub-circular; central area spanned by a fine proximal net and 2, broad conjunct transverse bars that taper towards the centre. Only the tranverse bars are visible in LM, they extend from the proximal surface to the distal surface.

Size: Type material length: 6.5-9.0 microns.


Phylogenetic relations: The evolutionary transition from T. eminens to T. occultatus is described in detail by Bybell & Self-Trail (1995), and more briefly by Romein (1979).

Most likely ancestor: Toweius eminens - at confidence level 4 (out of 5). Data source: Bybell & Self-Trail (1995, p.11, fig. 23).

Search data:
TagsLITHS: placolith, elliptical, CA: ca_conjunct, pores, bar,
CSPH: equant, monomorphic, CROSS-POLARS: rim-bicyclic, V-prominent, R-prominent,
MetricsLith size: 4->8µm;
The morphological data given here can be used on the advanced search page. See also these notes

Geological Range:
Last occurrence (top): within NP13 zone (49.11-50.50Ma, top in Ypresian stage). Data source: Perch-Nielsen 1985b
First occurrence (base): within NP9 zone (55.86-57.21Ma, base in Thanetian stage). Data source: Perch-Nielsen 1985

Plot of occurrence data:

References:

Bown, P. R. (2005d). Palaeogene calcareous nannofossils from the Kilwa and Lindi areas of coastal Tanzania (Tanzania Drilling Project 2003-4). Journal of Nannoplankton Research. 27(1): 21-95. gs V O

Bown, P. R. (2010). Calcareous nannofossils from the Paleocene/Eocene Thermal Maximum interval of southern Tanzania (TDP Site 14). Journal of Nannoplankton Research. 31(1): 11-38. gs V O

Bybell, L. M. & Self-Trail, J. (1995). Evolutionary, biostratigraphic and taxonomic study of calcareous nanofossils from a continuous Palaeocene-Eocene boundary section in New Jersey. U.S. Geological Survey, Professional Paper. 1554: 1-36. gs V O

Locker, S. (1967c). Neue, statigraphisch wichtige Coccolithophoriden (Flagellata aus dem norddeutschen Altetiar). Monatsberichte der Deutschen Akademie der Wissenschaften zu Berlin. 9(9-10): 758-769. gs

Perch-Nielsen, K. (1971c). Elektronenmikroskopische untersuchungen an Coccolithen und verwandten Formen aus dem Eozan von Danemark. Biologiske Skrifter, Kongelige Danske Videnskabernes Selskab. 18(3): 1-76. gs

Perch-Nielsen, K. (1985a). Cenozoic calcareous nannofossils. In, Bolli, H. M., Saunders, J. B. & Perch-Nielsen, K. (eds) Plankton Stratigraphy. Cambridge University Press, Cambridge 427-555. gs

Self-Trail, J. (2011). Paleogene Calcareous Nannofossils of the South Dover Bridge core, Southern Maryland (USA). Journal of Nannoplankton Research. 32(1): 1-28. gs V O

Shepherd, C. L. & Kulhanek, D. K. (2016). Eocene nannofossil biostratigraphy of the mid-Waipara River section, Canterbury Basin, New Zealand. Journal of Nannoplankton Research. 26(1): 33-59. gs V O

Varol, O. (1998). Palaeogene. In, Bown, P. R. (ed.) Calcareous Nannofossil Biostratigraphy. British Micropalaeontological Society Publication Series. 200-224. gs


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Toweius occultatus compiled by Jeremy R. Young, Paul R. Bown, Jacqueline A. Lees viewed: 14-4-2021

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Short stable page link: https://mikrotax.org/Nannotax3/index.php?id=1418 Go to Archive.is to create a permanent copy of this page - citation notes



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