Toweius pertusus


Classification: ntax_cenozoic -> Isochrysidales -> Prinsiaceae -> Toweius -> Toweius pertusus
Sister taxa: T. serotinus, T. pertusus, T. tovae, T. eminens, T. occultatus ⟩⟨ T. reticulum, T. selandianus, T. rotundus, T. patellus, T. callosus, T. magnicrassus ⟩⟨ T. gammation ⟩⟨ T. sp.

Taxonomy:

Citation: Toweius pertusus (Sullivan, 1965) Romein, 1979
Rank: Species
Basionym: Coccolithus pertusus Sullivan, 1965
Synonyms: Toweius craticulus Hay & Mohler 1967

Distinguishing features: Small to medium-sized (4-6 µm) with >=10 peforations.

Farinacci & Howe catalog pages: C. pertusus * , T. craticulus *


Morphology: Small to medium-sized; elliptical to sub-circular; central area with perforate net; perforations typically small and may be difficult to resolve in LM.


Phylogenetic relations: This is the main mid Palaeocene Toweius species and gives rise to larger species in the late Palaeocene and Early Eocene

Search data:
TagsLITHS: placolith, circular, elliptical, CA: ca_conjunct, grill, pores,
CSPH: equant, monomorphic, CROSS-POLARS: rim-bicyclic, V-prominent, R-prominent,
MetricsLith size: 3->8µm;
The morphological data given here can be used on the advanced search page. See also these notes

Geological Range:
Notes: Monechi et al. (2013) reccomend base of T. pertusus as alternative marker for NP2/3 boundary
Last occurrence (top): within NP14 zone (46.29-49.11Ma, top in Lutetian stage). Data source: [PRB rough estimate]
First occurrence (base): at base of NP4 zone (0% up, 63.3Ma, in Danian stage). Data source: Monechi et al. 2013, Fig 5

Plot of occurrence data:

References:

Agnini, C., Fornaciari, E. & Raffi, I. (2008). Three new species of calcareous nannofossil from Late Palaeocene and Early Eocene assemblages (Ocean Drilling Program Site 1262, Walvis Ridge, SE Atlantic Ocean). Journal of Nannoplankton Research. 30(1): 51-56. gs V O

Bown, P. R. & Dunkley Jones, T. (2012). Calcareous nannofossils from the Paleogene equatorial Pacific (IODP Expedition 320 Sites U1331-1334). Journal of Nannoplankton Research. 32(2): 3-51. gs V O

Bown, P. R. & Newsam, C. (2017). Calcareous nannofossils from the Eocene North Atlantic Ocean (IODP Expedition 342 Sites U1403–1411). Journal of Nannoplankton Research. 37(1): 25-60. gs V O

Bown, P. R. (2005d). Palaeogene calcareous nannofossils from the Kilwa and Lindi areas of coastal Tanzania (Tanzania Drilling Project 2003-4). Journal of Nannoplankton Research. 27(1): 21-95. gs V O

Bown, P. R. (2010). Calcareous nannofossils from the Paleocene/Eocene Thermal Maximum interval of southern Tanzania (TDP Site 14). Journal of Nannoplankton Research. 31(1): 11-38. gs V O

Bown, P. R. (2016). Paleocene calcareous nannofossils from Tanzania (TDP sites 19, 27 and 38). Journal of Nannoplankton Research. 36(1): 1-32. gs V O

Bown, P. R., Gibbs, S. J., Sheward, R., O’Dea, S. & Higgins, D. (2014). Searching for cells: the potential of fossil coccospheres in coccolithophore research. Journal of Nannoplankton Research. 34(special): 5-21. gs V O

Bybell, L. M. & Self-Trail, J. (1995). Evolutionary, biostratigraphic and taxonomic study of calcareous nanofossils from a continuous Palaeocene-Eocene boundary section in New Jersey. U.S. Geological Survey, Professional Paper. 1554: 1-36. gs V O

Hay, W. W. & Mohler, H. P. (1967). Calcareous nannoplankton from Early Tertiary rocks at Point Labau, France and Paleocene-Early Eocene correlations. Journal of Paleontology. 41(6): 1505-1541. gs

MacKenzie, D. T. & Wise,S. W. (1983). Paleocene and Eocene calcareous nannofossils from Deep Sea Drilling Project Legs 25 and 40, South and East of Africa. Initial Reports of the Deep Sea Drilling Project. 71(2): 1141-1170. gs V O

Mai, H., Hillebrandt-Habel, T., von Salis Perch-Nielsen, K. & Willems, H. (1998). Paleocene coccospheres from DSDP Leg 39, Site 356, Sao Paulo Plateau, S. Atlantic Ocean. Journal of Nannoplankton Research. 20(1): 21-29. gs V O

Monechi, S., Reale, V., Bernaola, G. & Balestra, B. (2013). The Danian/Selandian boundary at Site 1262 (South Atlantic) and in the Tethyan region: Biomagnetostratigraphy, evolutionary trends in fasciculiths and environmental effects of the Latest Danian Event. Marine Micropaleontology. 98(1): 28-40. gs

Okada, H. & Thierstein, H. R. (1979). Calcareous nannoplankton - Leg 43, Deep Sea Drilling Project. Initial Reports of the Deep Sea Drilling Project. 43: 507-573. gs V O

Perch-Nielsen, K. (1971b). Einige neue Coccolithen aus dem Paläozän der Bucht von Biskaya und dem Eozän der Labrador See. Bulletin of the Geological Society of Denmark. 21: 347-361. gs V O

Perch-Nielsen, K. (1971c). Elektronenmikroskopische untersuchungen an Coccolithen und verwandten Formen aus dem Eozan von Danemark. Biologiske Skrifter, Kongelige Danske Videnskabernes Selskab. 18(3): 1-76. gs

Perch-Nielsen, K. (1977a). Albian to Pleistocene calcareous nannofossils from the Western South Atlantic, DSDP Leg 39. Initial Reports of the Deep Sea Drilling Project. 39: 699-823. gs V O

Pospichal, J. J. & Wise, S. W. (1990c). Paleocene to Middle Eocene calcareous nannofossils of ODP sites 689 and 690, Maud Rise, Weddell Sea. Proceedings of the Ocean Drilling Program, Scientific Results. 113: 613-638. gs V O

Romein, A. J. T. (1979). Lineages in Early Paleogene calcareous nannoplankton. Utrecht Micropaleontological Bulletin. 22: 1-231. gs V O

Self-Trail, J. (2011). Paleogene Calcareous Nannofossils of the South Dover Bridge core, Southern Maryland (USA). Journal of Nannoplankton Research. 32(1): 1-28. gs V O

Shepherd, C. L. & Kulhanek, D. K. (2016). Eocene nannofossil biostratigraphy of the mid-Waipara River section, Canterbury Basin, New Zealand. Journal of Nannoplankton Research. 26(1): 33-59. gs V O

Sullivan, F. R. (1965). Lower Tertiary nannoplankton from the California Coast Ranges. II. Eocene. University of California Publications in Geological Sciences. 53: 1-74. gs

Thomsen, E. & Heilmann-Clausen, C. (1984). The Danian-Selandian boundary at Svejstrup with remarks on the biostratigraphy of the boundary in Western Denmark. Bulletin of the Geological Society of Denmark. 33: 341-362. gs V O

Varol, O. (1998). Palaeogene. In, Bown, P. R. (ed.) Calcareous Nannofossil Biostratigraphy. British Micropalaeontological Society Publication Series. 200-224. gs

Wise, S. W. & Wind, F. H. (1977). Mesozoic and Cenozoic calcareous nannofossils recovered by DSDP Leg 36 drilling on the Falkland Plateau, south-west Atlantic sector of the Southern Ocean. Initial Reports of the Deep Sea Drilling Project. 36(269-491): -. gs V O


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Toweius pertusus compiled by Jeremy R. Young, Paul R. Bown, Jacqueline A. Lees viewed: 14-4-2021

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