Distinguishing features:
Parent taxon (R. umbilicus group): Paleogene elliptical reticulofenestrids with open central area (spanned by delicate proximal net, not visible in lm)
This taxon: Very large (>14 µm) elliptical reticulofenestrids, with delicate net
| LITHS: placolith, elliptical, CA: ca_conjunct, grill, vacant, CSPH: equant, monomorphic, CROSS-POLARS: rim-unicyclic, R-prominent, |
| Lith size: 14->20µm; Data source notes: Size range from Backman & Hermelin 1986 |
Geological Range:
Notes: The first appearance of very large, >14μm, elliptical reticulofenestrids (i.e. R. umbilicus) has long been used as a distinctive stratigraphic marker (e.g. Backman & Hermelin, 1986). In the Exp. 342 material these sizes are attained by subcircular to elliptical reticulofenestrids (similar to R. wadeae and R. umbilicus) in Subzone NP15b, around 3 m.y. prior to the level typically cited for this biohorizon. Agnini et al. (2014) suggest that the use of the first common occurrence of R. umbilicus may represent a more consistent horizon, above the last occurrence of C. gigas. [Bown & Newsam 2017]
Last occurrence (top): at top of NP22 zone (100% up, 32Ma, in Rupelian stage). Data source: zonal marker (see Perch-Nielsen 1985)
First occurrence (base): within NP16 zone (40.40-42.87Ma, base in Lutetian stage). Data source: Perch-Nielsen 1985
Plot of occurrence data:
Agnini, C., et al. (2014). Biozonation and biochronology of Paleogene calcareous nannofossils from low and middle latitudes. Newsletters on Stratigraphy. 47: 131-181. gs Backman, J. & Hermelin, J. O. R. (1986). Morphometry of the Eocene nannofossil Reticulofenestra umbilicus lineage and its biochronological consequences. Palaeogeography Palaeoclimatology Palaeoecology. 57(1): 103-116. gs Bown, P. R. & Dunkley Jones, T. (2012). Calcareous nannofossils from the Paleogene equatorial Pacific (IODP Expedition 320 Sites U1331-1334). Journal of Nannoplankton Research. 32(2): 3-51. gs O Bown, P. R. & Newsam, C. (2017). Calcareous nannofossils from the Eocene North Atlantic Ocean (IODP Expedition 342 Sites U1403–1411). Journal of Nannoplankton Research. 37(1): 25-60. gs O Bown, P. R. (2005d). Palaeogene calcareous nannofossils from the Kilwa and Lindi areas of coastal Tanzania (Tanzania Drilling Project 2003-4). Journal of Nannoplankton Research. 27(1): 21-95. gs O Bown, P. R., Gibbs, S. J., Sheward, R., O’Dea, S. & Higgins, D. (2014). Searching for cells: the potential of fossil coccospheres in coccolithophore research. Journal of Nannoplankton Research. 34(S): 5-21. gs Bybell, L. (1975). Middle Eocene calcareous nannofossils at Little Stave Creek, Alabama. Tulane Studies in Geology and Paleontology. 11(4): 177-247. gs O De Kaenel, E. & Mojon, P. -O. (2023). Calcareous nannofossil biostratigraphy of the Eocene/Oligocene boundary (Zones NP19–21) in the parautochthonous Subalpine UMM of the Swiss Molasse Basin (Canton Fribourg, Western Switzerland). Journal of Nannoplankton Research. 41(2): 64-96. gs O de Kaenel, E. & Villa, G. (1996). Oligocene-Miocene calcareous nannofossil biostratigraphy and paleoeecology from the Iberian Abyssal Plain. Proceedings of the Ocean Drilling Program, Scientific Results. 149: 79-145. gs O Dunkley Jones, T., Bown, P. R. & Pearson, P. (2009). Exceptionally well preserved upper Eocene to lower Oligocene calcareous nannofossils (Prymnesiophycidae) from the Pande Formation (Kilwa Group), Tanzania. Journal of Systematic Palaeontology. 7(4): 359-411. gs Fioroni, C., Villa, G., Persico, D., Wise, S. W. & Pea, L. (2012). Revised middle Eocene-upper Oligocene calcareous nannofossil biozonation for the Southern Ocean. Revue de Micropaléontologie. 55(2): 53-70. gs Fioroni, C., Villa, G., Persico, D. & Jovane, L. (2015). Middle Eocene-Lower Oligocene calcareous nannofossil biostratigraphy and paleoceanographic implications from Site 711 (equatorial Indian Ocean). Marine Micropaleontology. 118: 50-62. gs Haq, B. U. (1968). Studies on upper Eocene calcareous nannoplankton from NW Germany. Stockholm Contributions in Geology. 18: 13-74. gs O Jones, A. P. & Dunkley Jones, T. (2020). Middle Eocene to Early Oligocene calcareous nannofossils from the Nanggulan Formation, Java, Indonesia. Journal of Nannoplankton Research. 38(1): 57-79. gs Kamptner, E. (1948). Coccolithen aus dem Torton des Inneralpinen Wiener Beckens. Anzeiger der Österreichische Akademie der Wissenschaften, Wien. 157: 1-16. gs Levin, H. L. (1965). Coccolithophoridae and related microfossils from the Yazoo Formation (Eocene) of Mississippi. Journal of Paleontology. 39(2): 265-272. gs Martini, E. & Ritzkowski, S. (1968). Was ist das 'Unter-Oligozän'? Eine Analyse der Beyrich'schen und v Koenen'schen Fassung der Stufe mit Hilfe des fossilen Nannoplanktons. Nachrichten der Akademie der Wissenschaften in Gottingen, II Mathematisch-Physikalische Klasse. 1968, 13: 231-250. gs Müller, C. (1979). Calcareous nannofssils from the North Atlantic (Leg 48). Initial Reports of the Deep Sea Drilling Project. 48: 589-639. gs Perch-Nielsen, K. (1971c). Elektronenmikroskopische untersuchungen an Coccolithen und verwandten Formen aus dem Eozan von Danemark. Biologiske Skrifter, Kongelige Danske Videnskabernes Selskab. 18(3): 1-76. gs O Perch-Nielsen, K. (1972). Remarks on Late Cretaceous to Pleistocene coccoliths from the North Atlantic. Initial Reports of the Deep Sea Drilling Project. 12: 1003-1069. gs Perch-Nielsen, K. (1985). Cenozoic calcareous nannofossils. In, Bolli, H. M., Saunders, J. B. & Perch-Nielsen, K. (eds) Plankton Stratigraphy. Cambridge University Press, Cambridge (1): 427-555. gs Self-Trail, J. (2011). Paleogene Calcareous Nannofossils of the South Dover Bridge core, Southern Maryland (USA). Journal of Nannoplankton Research. 32(1): 1-28. gs Shepherd, C. L. & Kulhanek, D. K. (2016). Eocene nannofossil biostratigraphy of the mid-Waipara River section, Canterbury Basin, New Zealand. Journal of Nannoplankton Research. 36(1): 33-59. gs O Varol, O. (1998). Palaeogene. In, Bown, P. R. (ed.) Calcareous Nannofossil Biostratigraphy. British Micropalaeontological Society Publication Series . 200-224. gs Wei, W. & Wise, S. W. (1990). Middle Eocene to Pleistocene calcareous nannofossils recovered by Ocean Drilling Program leg 113 in the Weddell Sea. Proceedings of the Ocean Drilling Program, Scientific Results. 113: 639-666. gs Wise, S. W. (1983). Mesozoic and Cenozoic calcareous nannofossils recovered by DSDP Leg 71 in the Falkland Plateau region, Southwest Atlantic Ocean. Initial Reports of the Deep Sea Drilling Project. 71: 481-550. gs OReferences:

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Reticulofenestra umbilicus compiled by Jeremy R. Young, Paul R. Bown, Jacqueline A. Lees viewed: 22-7-2026
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