bforams@mikrotax - Ammolagena clavata

NB This site is under active development, current content in the main database is primarily from Bolli et al 1994, Hayward et al. 2012, Holbourn et al. 2013, Holbourn and Kaminski 1997, Alegret & Thomas 2001, and Katz et al. 2003. It is intended to expand and develop this to provide rather comprehensive coverage of Mesozoic and Cenozoic deep-sea benthic foraminifera.

Taxonomy

Citation: Ammolagena clavata (Jones & Parker), 1860

Basionym: Trochammina irregularis var. clavata Jones & Parker, 1860

Synonyms:

Trochammina irregularis (d’Orbigny) var. clavata Jones and Parker. –Carpenter, Parker, and Jones, 1862, p. 142, pl. 11, fig. 6. [according to Holbourn et al. 2013]
Ammolagena clavata (Jones and Parker). –Loeblich and Tappan, 1987, p. 49, pl. 36, fig. 16. –Zheng, 1988, p. 39, pl. 6, fig. 6. –Kaminski et al., 1990, p. 365, pl. 3, fig. 1. –Charnock and Jones, 1990, p. 155, pl. 2, fig. 8; pl. 14, fig. 6. –Jones, 1994, p. 46, pl. 41, figs. 12–16. –Loeblich and Tappan, 1994, p. 14, pl. 4, figs. 1–4. –Holbourn and Kaminski, 1997, p. 36, pl. 5, fig. 6. –Kaminski and Gradstein, 2005, pp. 165–168, fig. 21, pl. 21. –Milker and Schmiedl, 2012, p. 28, fig. 9.5. [according to Holbourn et al. 2013]

Taxonomic discussion: Type species of the genus Ammolagena Eimer and Fickert, 1899. Test is often found attached to shell fragments or to other foraminiferal tests. Common in Upper Cretaceous to Paleogene flysch-type assemblages. [Holbourn et al. 2013]
According to Kaminski & Gradstein (2005) the existing definition of Ammolagena Eimer and Fickert, 1899 needs to be emended to take into account the presence of a supplementary aperture. The original description of Jones & Parker does not mention the presence of a second aperture at the base of the proloculus, nor is this feature mentioned by Loeblich & Tappan (1987). The specimen figured by Loeblich & Tappan (1964) from ALBATROSS Sta. D2385 in the Gulf of Mexico (USNM 433468) does not have a secondary aperture. However, many Recent specimens from ALBATROSS stations in the USNM collections display secondary apertures which are often surrounded by a lip. Wright's (1891) colour drawing of a specimen collected in 750 fathoms off SW Ireland clearly shows a small, round second aperture surrounded by a distinct lip, and Rhumbler's (1913) colour drawing shows a second aperture which is clearly labelled. Cushman (1918) noted that both microsphaeric and megalosphaeric individuals occur, with microsphaeric specimens having a smaller proloculus and longer tubular chamber. Collins (1958) found a specimen that has a second pyriform chamber arising from the tubular extension of the first and noted "the definition of the genus should be emended to include the multilocular condition".
Pflum & Frerichs (1976) reported that A. clavata has a depressed upper depth limit near the mouth of the Mississippi River. In this respect, A. clavata is a species which is environmentally sensitive and adversely affected by river discharge or "deltaic influence". Schröder-Adams & McNeil (1994) recorded it in low numbers in the deltaic shelf facies of the Kugmallit sequences of the Beaufort-MacKenzie Basin.

Description

Test forms a large, ovoid proloculus followed by a narrow, tubular chamber. Chamber walls are non-calcareous, finely agglutinated, and smooth. The aperture is a terminal and round opening at the end of the tube. A secondary aperture may also be present at the base of the proloculus. [Holbourn et al. 2013]

The lectotype is 0.27 mm wide across the proloculus. Paralectotypes from the Mediterranean are up to 1.9 mm in length. The width of the proloculus varies from 0.20 to 0.34 mm.[Kaminski 7 Gradstein (2005)]

Biogeography and Palaeobiology

Worldwide. See also Observed Occurrences in Kaminski and Gradstein (2005).

Bathyal and abyssal. [Holbourn et al. 2013]

Biostratigraphic distribution

Geological Range:
Notes: Early Cretaceous to Recent.
Last occurrence (top): Extant. Data source: Holbourn et al. 2013
First occurrence (base): within Early Cretaceous Epoch (100.50-143.10Ma, base in Berriasian stage). Data source: Holbourn et al. 2013

Plot of occurrence data:
NB The ranges given for genera and higher taxa are the total ranges of species included in that taxon in this database - so the true total range of the taxon will usually be longer

References:

Brady, H. B. (1884). Report on the foraminifera dredged by H.M.S. Challenger during the years 1873–1876. Report on the Scientific Results of the Voyage of H.M.S. Challenger during the years 1873-1876. 9: 1-814. gs

Carpenter, W. B., Parker, W. K. & Jones, J. P. (1862). Introduction to the Study of the Foraminifera. Ray Society, London. 1-319. gs

Charnock, M. A. & Jones, R. W. (1990). Agglutinated foraminifera from the Palaeogene of the North Sea. In, Hemleben, C., Kaminski, M. A., Kuhnt, W. , & Scott, D. B. (eds) Paleoecology, Biostratigraphy, Paleoceanography and Taxonomy of Agglutinated Foraminifera,. Nato ASI (Advanced Study Institute) Series . 139-244. gs

Cushman, J. A. (1946b). Upper Cretaceous foraminifera of the Gulf coastal region of the United States and adjacent areas. U.S. Geological Survey, Professional Paper. 206: pp 1-241-. gs

Holbourn, A. E. L. & Kaminski, M. A. (1997). Lower Cretaceous benthic foraminifera of the Indian Ocean. Grzybowski Foundation Special Publication. 4: 1-172. gs

Holbourn, A., Henderson, A. S. & MacLeod, N. (2013). Atlas of Benthic Foraminifera. Natural History Museum, London. 1-651. gs

Jones, J. P. & Parker, W. K. (1860). On the Rhizopodal fauna of the Mediterranean compared with that of the Italian and some other Tertiary deposits. Quarterly Journal of the Geological Society of London. 16: 292-307. gs

Jones, R. W. (1994). The Challenger Foraminifera. Oxford University Press, Oxford. 1-149. gs

Kaminski, M. A. & Gradstein, F. M. (2005). Atlas of Palaeogene cosmopolitan deep-water agglutinated foraminifera. Grzybowski Foundation Special Publication. (10): 1-547. gs

Kaminski, M. A., Gradstein, F. M., Goll, R. M. & Greig, D. (1990). Biostratigraphy and Paleoecology of deep-water aggutinated foraminifera at ODP Site 643, Norwegian–Greenland Sea. In, Hemleben, C., Kaminski, M. A., Kuhnt, W. , & Scott, D. B. (eds) Paleoecology, Biostratigraphy, Paleoceanography and Taxonomy of Agglutinated Foraminifera,. Nato ASI (Advanced Study Institute) Series . 327: 345-386. gs

Loeblich, A. R. Jr & Tappan, H. (1987). Foraminiferal Genera and their Classification. Van Nostrand Reinhold, New York. 1-970. gs

Loeblich, A. & Tappan, H. (1994). Foraminifera of the Sahul shelf and Timor Sea. Cushman Foundation for Foraminiferal Research, Special Publication. 31: 1-661. gs

Milker, Y. & Schmiedl, G. (2012). A taxonomic guide to modern benthic shelf foraminifera of the western Mediterranean Sea. Palaeontologia Electronica. 15(2;16A ): 1-134. gs

Zheng, S. Y. (1988). The Agglutinated and Porcelaneous Foraminifera of the East China Sea. China Ocean Press, Beijing. 1-337. gs

Ammolagena clavata

Taxonomy

Description

Biogeography and Palaeobiology

Biostratigraphic distribution

References:

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