Acarininasoldadoensisangulosa (Bolli).— Hillebrandt, 1976:345, pl. 5: fig. 11 [Zone P9, Agost, Alicante Province, Spain].—Fleisher, 1974:1014, pl. 4: fig. 1 [lower Eocene Zone P9, DSDP Site 220, Arabian Sea].
?Turborotalia (Acarinina) soldadoensisangulosa (Bolli).— Samuel and others, 1972:190, pl. 68, figs. 1a-c [lower Eocene Globorotaliasubbotinae Zone, Hradisko, Middele Va_h Valley, West Carpathian].
Muricoglobigerinasoldadoensisangulosa (Bolli).—Blow, 1979:1122, pl. 109: fig. 9, pl. 131: figs. 4-5 [lower Eocene Zone P6 of Blow, 1979), DSDP Hole 47.2, Shatsky Rise, north-west Pacific Ocean]—?Belford, 1984:30, pl. 22: figs. 9-12 [lower Eocene Zones P8-9, Papua New Guinea].
Taxonomic discussion: This distinctive taxon is a common component of lower Eocene (sub)tropical assemblages where it is associated with other acarininids (soldadoensis, interposita, pentacamerata, pseudotopilensis) and morozovellids (subbotinae, formosa, gracilis, aragonensis). In the course of our studies in Egypt we have found that this taxon has its FAD just below the base of the CIE / PETM interval, i.e., within the uppermost part of Zone P5 (as redefined in Berggren and Pearson, 2005). [Berggren et al. 2006]
Distinguishing features: Parent taxon (Acarinina): Moderate to low trochospire; chambers ovoid, usually 4-6 in final whorl. Wall muricate with pustules on umbilical shoulders; This taxon: Last whorl with strongly angular disposition of the chambers; outline strongly lobulate.
NB These concise distinguishing features statements are used in the tables of daughter-taxa to act as quick summaries of the differences between e.g. species of one genus. They are being edited as the site is developed and comments on them are especially welcome.
Description
Diagnostic characters: The diagnostic features of Acarininaangulosa are the strongly angular disposition of the chambers of the last whorl and the strongly lobulate peripheral outline. [Berggren et al. 2006] Morphology: Low trochospiral test with lobulate peripheral outline; about 10-12 chambers in 2½ whorls; 4½-5 subangular chambers in last whorl gradually increasing in size, separated by straight, radial and depressed sutures; spiral side flat to slightly convex, sutures curved/oblique; umbilicus moderately wide, open, deep; aperture a low interiomarginal-umbilical arch extending nearly to the periphery; axial/edge view subangular. [Berggren et al. 2006] Wall type:
Soldan et al 2014 f05-4d.JPG
Soldan et al 2014 f05-4e.JPG
Muricate, normal perforate, nonspinose. [Berggren et al. 2006] Size: Maximum diameter of holotype 0.40mm, thickness 0.28 mm. [Berggren et al. 2006]
Character matrix
test outline:
Lobate
chamber arrangement:
Trochospiral
edge view:
Inequally biconvex
aperture:
Umbilical-extraumbilical
sp chamber shape:
Inflated
coiling axis:
Low
periphery:
N/A
aperture border:
N/A
umb chbr shape:
Inflated
umbilicus:
Wide
periph margin shape:
Subangular
accessory apertures:
None
spiral sutures:
Strongly depressed
umb depth:
Deep
wall texture:
Moderately muricate
shell porosity:
Finely Perforate: 1-2.5µm
umbilical or test sutures:
Strongly depressed
final-whorl chambers:
4.5-5
N.B. These characters are used for advanced search. N/A - not applicable
Biogeography and Palaeobiology
Geographic distributionRanges from tropical to temperate regions; Arabian Sea, north Pacific Ocean (DSDP Site 47), New Jersey Coastal Plain (Bass River), Tethyan deposits of northern Africa (Egypt) and occurrences in Spain (Alicante Province) and New Guinea. [Berggren et al. 2006]
Aze et al. 2011 summary: Low to middle latitudes; based on Berggren et al. (2006b) Isotope paleobiologyNo data available. [Berggren et al. 2006] Aze et al. 2011 ecogroup 1 - Open ocean mixed-layer tropical/subtropical, with symbionts; based on comparison with other species of the genus. Phylogenetic relationsThis species evolved from Acarininasoldadoensis in uppermost Zone P5 and probably gave rise to A. cuneicamerata. [Berggren et al. 2006]
Geological Range: Notes: Zone P5 (uppermost part) to Zone E7. Note that the Paleocene part of the range was not recorded by Olsson and others (1999). [Berggren et al. 2006] Last occurrence (top): within E7 zone (45.72-50.20Ma, top in Lutetian stage). Data source: Eocene Atlas First occurrence (base): in mid part of P5 zone (50% up, 56.5Ma, in Thanetian stage). Data source: Eocene Atlas
Plot of occurrence data:
Range-bar - range as quoted above, pink interval top occurs in, green interval base occurs in.
Triangles indicate an event for which a precise placement has been suggested.
Grey shading between taxa indicates intergrading species within an anagenetic lineage (data from
Lamyman et al. 2026).
(NB There is no histogram as there are no occurrence records for the taxon in the Neptune database) Parent: Acarinina
Primary source for this page: Berggren et al. 2006 - Eocene Atlas, chap. 9, p. 263
References:
Belford, D. J. (1984). Tertiary foraminifera and age of sediments, Ok Tedi-Wabag, Papua New Guinea. Australia Bureau of Mineral Resources Geology and Geophysics, Bulletin. 216: 1-52. gs
Berggren, W. A. & Pearson, P. N. (2005). A revised tropical to subtropical Paleogene planktonic foraminiferal zonation. Journal of Foraminiferal Research. -. gs
Berggren, W. A., Pearson, P. N., Huber, B. T. & Wade, B. S. (2006b). Taxonomy, biostratigraphy, and phylogeny of Eocene Acarinina. In, Pearson, P. N., Olsson, R. K., Hemleben, C., Huber, B. T. & Berggren, W. A. (eds) Atlas of Eocene Planktonic Foraminifera. Cushman Foundation for Foraminiferal Research, Special Publication . 41(Chap 9): 257-326. gsO
Blow, W. H. (1979). The Cainozoic Globigerinida: A study of the morphology, taxonomy, evolutionary relationships and stratigraphical distribution of some Globigerinida (mainly Globigerinacea). E. J. Brill, Leiden. 2: 1-1413. gs
Bolli, H. M. (1957d). The genera Globigerina and Globorotalia in the Paleocene-Lower Eocene Lizard Springs Formation of Trinidad. In, Loeblich, A. R. , Jr., Tappan, H., Beckmann, J. P., Bolli, H. M., Montanaro Gallitelli, E. & Troelsen, J. C. (eds) Studies in Foraminifera. U.S. National Museum Bulletin . 215: 61-82. gs
Bolli, H. M. (1957a). Planktonic foraminifera from the Eocene Navet and San Fernando formations of Trinidad. In, Loeblich, A. R. , Jr., Tappan, H., Beckmann, J. P., Bolli, H. M., Montanaro Gallitelli, E. & Troelsen, J. C. (eds) Studies in Foraminifera. U.S. National Museum Bulletin . 215: 155-172. gs
Fleisher, R. L. (1974a). Cenozoic planktonic foraminifera and biostratigraphy, Arabian Sea, Deep Sea Drilling Project, Leg 23A. Initial Reports of the Deep Sea Drilling Project. 23: 1001-1072. gsO
Hillebrandt, A. , von (1976). Los foraminiferos planctonicos, nummulitidos y coccolitoforidos de la zona de Globorotalia palmerae del Cuisiense (Eoceno inferior) en el SE de Espana, (Provincias de Murcia y Alicante. Revista Española de Micropaleontología. 8(3): 323-394. gsO
Samuel, O., Borza, K. & Kohler, E. (1972). Microfauna and lithostratigraphy of the Paleogene and adjacent Cretaceous of the Middle Vah Valley (West Carpathian). Geologicky Ustav Dionyza Stura, Bratislava. 1-264. gs
Soldan, D. M., Petrizzo, M. R. & Silva, I. P. (2014). Pearsonites, a new Paleogene planktonic foraminiferal genus for the broedermanni lineage. Journal of Foraminiferal Research. 44: 17-27. gs
Acarinina angulosa compiled by the pforams@mikrotax project teamviewed: 19-8-2026