Muricoglobigerinaesnehensis (Nakkady).—Pearson and others, 1993: pl. 1: fig. 20 [middle Eocene Zone P11-P12, DSDP Site 523, Walvis Ridge, South Atlantic Ocean]. [Not Nakkady, 1950.]
Acarininaaquiensis (Loeblich and Tappan).—Lu and Keller, 1995: pl. 2: figs. 24-26 [Lower Eocene Zone P9, DSDP Site 577, Shatsky Rise, north-west Pacific Ocean]. [Not Loeblich and Tappan, 1957.]
Acarinina cf. subsphaerica (Subbotina).—Pearson and others, 2004:37, pl. 2, fig. 4 [middle Eocene, Zone P11, Tanzania Drilling Project Site 2, Kilwa Masoko, Tanzania]. [Not Subbotina, 1947.]
Taxonomic discussion: Despite being a relatively common form in the lower and middle Eocene, Acarininapseudosubsphaerica n. sp. has not been formally recorded. Acarininaalticonica Fleisher is closely related and the two species have been observed to intergrade in Zone E6/E7 (undifferentiated) of Tanzania Drilling Project Site 2 (P .N. Pearson, unpublished data). Lu and Keller (1995) recorded a first occurrence in Zone P9 (=E7) of Acarininaaquiensis in DSDP Site 577 (Shatsky Rise, north-west Pacific Ocean), which is regarded as conspecific here, and not referable to Loeblich and Tappan’s species (which is a Paleocene form from the Aquia Formation of Virginia and Maryland probably related to/conspecific with Acarininasubsphaerica; see Berggren and Norris, 1997). [Berggren et al. 2006]
Distinguishing features: Parent taxon (Acarinina): Moderate to low trochospire; chambers ovoid, usually 4-6 in final whorl. Wall muricate with pustules on umbilical shoulders; This taxon: Like A. alticonica but higher-spired; chambers less closely appressed; and umbilicus broader.
NB These concise distinguishing features statements are used in the tables of daughter-taxa to act as quick summaries of the differences between e.g. species of one genus. They are being edited as the site is developed and comments on them are especially welcome.
Description
Diagnostic characters: Acarininapseudosubsphaerica n. sp. is distinguished from all other middle Eocene acarininids by its high spire, small size and subspherical shape. It is similar in morphology to the Paleocene species, A. subsphaerica (Subbotina), but differs from that form and A. alticonica Fleisher by having less closely appressed chambers and a broader umbilicus. [Berggren et al. 2006] Morphology: Compact subspherical test coiled in a high trochospire of about two whorls with circular, slightly lobate outline; 4-6 moderately inflated, radially compressed chambers in final whorl; dorsal sutures wide and depressed, between circular chambers; primary aperture slit-like or a low, broad arch, situated in a wide, open umbilicus, and often covered by a small, sparsely muricate final chamber.[Berggren et al. 2006] Wall type: Weakly muricate, cancellate on spiral side with large, bladed muricae on umbilical side, normal perforate, nonspinose.[Berggren et al. 2006]
Size: Maximum diameter of holotype 0.19 mm, thickness 0.16 mm.[Berggren et al. 2006]
Character matrix
test outline:
Subcircular
chamber arrangement:
Trochospiral
edge view:
Equally biconvex
aperture:
Umbilical
sp chamber shape:
Globular
coiling axis:
Moderate-high
periphery:
N/A
aperture border:
N/A
umb chbr shape:
Subtriangular
umbilicus:
Wide
periph margin shape:
Broadly rounded
accessory apertures:
None
spiral sutures:
Weakly depressed
umb depth:
Deep
wall texture:
Coarsely muricate
shell porosity:
Finely Perforate: 1-2.5µm
umbilical or test sutures:
Weakly depressed
final-whorl chambers:
4-6
N.B. These characters are used for advanced search. N/A - not applicable
Biogeography and Palaeobiology
Geographic distributionProbably widely distributed in tropical and mid-latitude sites. [Berggren et al. 2006]
Aze et al. 2011 summary: Low to middle latitudes; based on Berggren et al. (2006b) Isotope paleobiologyRecorded (under the name Muricoglobigerinaesnehensis) as a mixed layer dwelling species by Pearson and others, (1993). [Berggren et al. 2006] Aze et al. 2011 ecogroup 1 - Open ocean mixed-layer tropical/subtropical, with symbionts. Based on very heavy _13C and relatively light _18O. Sources cited by Aze et al. 2011 (appendix S3): Pearson et al. (1993) Phylogenetic relationsDescended from A. alticonica. [Berggren et al. 2006]
Geological Range: Notes: Zone E7 to Zone E10 [Berggren et al. 2006] Last occurrence (top): within E10 zone (41.89-43.23Ma, top in Lutetian stage). Data source: Eocene Atlas First occurrence (base): within E7a subzone (48.31-50.20Ma, base in Ypresian stage). Data source: Eocene Atlas
Plot of occurrence data:
Range-bar - range as quoted above, pink interval top occurs in, green interval base occurs in.
Triangles indicate an event for which a precise placement has been suggested
Histogram - Neptune occurrence data from DSDP and ODP proceedings. Pale shading <50 samples in time bin. Interpret with caution & read these notes
Taxon plotted: Acarinina pseudosubsphaerica, synonyms included - Acarinina pseudosubsphaerica;
Primary source for this page: Berggren et al. 2006 - Eocene Atlas, chap. 9, p. 303
References:
Berggren, W. A. & Norris, R. D. (1997). Biostratigraphy, phylogeny and systematics of Paleocene trochospiral planktonic foraminifera. Micropaleontology. 43(supplement 1): 1-116. gs
Berggren, W. A., Pearson, P. N., Huber, B. T. & Wade, B. S. (2006b). Taxonomy, biostratigraphy, and phylogeny of Eocene Acarinina. In, Pearson, P. N., Olsson, R. K., Hemleben, C., Huber, B. T. & Berggren, W. A. (eds) Atlas of Eocene Planktonic Foraminifera. Cushman Foundation for Foraminiferal Research, Special Publication . 41(Chap 9): 257-326. gsO
Loeblich, A. R. & Tappan, H. (1957b). Planktonic foraminifera of Paleocene and early Eocene Age from the Gulf and Atlantic coastal plains. In, Loeblich, A. R. , Jr., Tappan, H., Beckmann, J. P., Bolli, H. M., Montanaro Gallitelli, E. & Troelsen, J. C. (eds) Studies in Foraminifera. U.S. National Museum Bulletin . 215: 173-198. gs
Lu, G. & Keller, G. (1995). Planktic foraminiferal faunal turnovers in the subtropical Pacific during the Late Paleocene to Early Eocene. Journal of Foraminiferal Research. 25: 97-116. gs
Nakkady, S. E. (1950). A new foraminiferal fauna from the Esna shales and Upper Cretaceous chalk of Egypt. Journal of Paleontology. 24(6): 675-692. gs
Pearson, P. N. & Berggren, W. A. (2006). Taxonomy, biostratigraphy, and phylogeny of Morozovelloides n. gen. In, Pearson, P. N., Olsson, R. K., Hemleben, C., Huber, B. T. & Berggren, W. A. (eds) Atlas of Eocene Planktonic Foraminifera. Cushman Foundation for Foraminiferal Research, Special Publication . 41(Chap 10): 327-342. gsO
Pearson, P. N., Shackleton, N. J. & Hall, M. A. (1993). Stable isotope paleoecology of middle Eocene planktonic foraminifera and multi-species isotope stratigraphy, DSDP Site 523, South Atlantic. Journal of Foraminiferal Research. 23: 123-140. gs
Acarinina pseudosubsphaerica compiled by the pforams@mikrotax project teamviewed: 25-4-2025