Acarininaquetra (Bolli).—Fleisher, 1974, pl. 3: fig. 3 [Zone P8, DSDP Site 220, Arabian Sea].—Berggren, 1977: 251, chart no. 10 (reillustrations from literature).—Warraich and others, 2000:193, fig. 17.13-15 [Zone P7, Rakhi Nala section, Sulaiman Range, Pakistan].—Warraich and Ogasawara, 2001:29, figs. 7.4-6 [Zone P7, Rakhi Nala section, Sulaiman Range, Pakistan].
Globorotalia (Truncorotaloides) quetra (Bolli).—Blow, 1979:1034-1036 (note p. 1036 misprinted as 1136), pl. 122: figs. 4, 7-9 and pl. 200: figs. 5, 6 ; pl. 123: figs. 6-9, pl. 202: figs. 1-6 [lower Eocene Zone P7, DSDP Site 47, Shatsky Rise, north-west Pacific Ocean]; pl. 129: fig. 5 [lower Eocene Zone P8a, DSDP Site 47, Shatsky Rise, northwest Pacific Ocean]; pl. 132: figs. 7-9 and pl. 201: figs. 4-6 [lower Eocene Zone P7, DSDP Site 47, Shatsky Rise, northwest Pacific Ocean]; pl. 140: fig. 4 [lower Eocene Zone P8b, DSDP Site 20, South Atlantic Ocean].
Morozovellaquetra (Bolli).—Snyder and Waters, 1985: 448, pl. 9: figs. 4-6 [lower Eocene Zone P6, DSDP Site 550, Porcupine Abyssal Plain, north-east Atlantic Ocean].
Taxonomic discussion: Acarininaquetra is a distinctive form that with its cuneate chambers and peripheral concentration of muricae, bears a superficial resemblance (in some instances) to the stratigraphically younger (middle Eocene) A. topilensis. [Berggren et al. 2006]
Distinguishing features: Parent taxon (Acarinina): Moderate to low trochospire; chambers ovoid, usually 4-6 in final whorl. Wall muricate with pustules on umbilical shoulders; This taxon: Like A. pseudotopilensis but with angular test; chambers loosely disposed to each other at ~ 90°; and with a distinct, but discontinuous, peripheral muricocarina.
NB These concise distinguishing features statements are used in the tables of daughter-taxa to act as quick summaries of the differences between e.g. species of one genus. They are being edited as the site is developed and comments on them are especially welcome.
Description
Diagnostic characters: This taxon is distinguished by its angular test whose chambers are loosely disposed sequentially to each other at ~ 90o and with a distinct, but discontinuous, peripheral muricocarina. [Berggren et al. 2006] Morphology: Low trochospiral; test subquadrate, planoconvex; peripheral margin lobulate; chambers on umbilical side moderately inflated, flattened on spiral side; 4 broadly subtriangular chambers visible in umbilical view; anterior and posterior margins of last chamber flattened resulting in disjunct geometry with bordering chambers and conical apex of last chamber; sutures depressed/incised, curved to sinuous, radial; umbilicus relatively wide, deep; aperture an umbilical-extraumbilical, arch extending towards (but not to) the peripheral margin; 10-12 chambers in 2½ to 3 whorls on spiral side; chambers loosely disposed, meeting almost at right angles, increasing gradually in size; final chamber sometimes reduced in size; chambers lens-shaped, tangentially longer than radially broad in general, overlapping; sutures curved; discrete intercameral openings visible on some individuals (depending on preservation); planoconvex in edge view, final chamber distinctly anguloconical with high conical angle; periphery distinctly, but discontinuously, muricocarinate [Berggren et al. 2006] Wall type: Moderately to strongly muricate with concentration of muricae along peripheral margin of last whorl, nonspinose, normal perforate. [Berggren et al. 2006] Size: Maximum diameter of holotype 0.50 mm, thickness 0.38 mm. [Berggren et al. 2006]
Character matrix
test outline:
Subquadrate
chamber arrangement:
Trochospiral
edge view:
Planoconvex
aperture:
Umbilical-extraumbilical
sp chamber shape:
Inflated
coiling axis:
Very low
periphery:
N/A
aperture border:
N/A
umb chbr shape:
Inflated
umbilicus:
Wide
periph margin shape:
Broadly rounded
accessory apertures:
Sutural
spiral sutures:
Moderately depressed
umb depth:
Deep
wall texture:
Coarsely muricate
shell porosity:
Finely Perforate: 1-2.5µm
umbilical or test sutures:
Moderately depressed
final-whorl chambers:
4-4
N.B. These characters are used for advanced search. N/A - not applicable
Biogeography and Palaeobiology
Geographic distributionWidely distributed in (sub)tropical regions (Caribbean, Atlantic, Indo-Pacific, Tethyan/Mediterranean regions, North Caucasus) [Berggren et al. 2006]
Aze et al. 2011 summary: Low latitudes; based on Berggren et al. (2006b) Isotope paleobiologyNo data available [Berggren et al. 2006] Aze et al. 2011 ecogroup 1 - Open ocean mixed-layer tropical/subtropical, with symbionts; based on comparison with other species of the genus. Phylogenetic relationsDescended from A. pseudotopilensis. [Berggren et al. 2006]
Most likely ancestor:Acarinina pseudotopilensis - at confidence level 4 (out of 5). Data source: Berggren et al. (2006) fig9.2.
Biostratigraphic distribution
Geological Range: Notes: Zone E3 (upper part) to Zone E6. [Berggren et al. 2006] Last occurrence (top): within E6 zone (50.20-50.67Ma, top in Ypresian stage). Data source: Eocene Atlas First occurrence (base): in upper part of E3 zone (80% up, 54.7Ma, in Ypresian stage). Data source: Eocene Atlas
Plot of occurrence data:
Range-bar - range as quoted above, pink interval top occurs in, green interval base occurs in.
Triangles indicate an event for which a precise placement has been suggested
Histogram - Neptune occurrence data from DSDP and ODP proceedings. Pale shading <50 samples in time bin. Interpret with caution & read these notes
Primary source for this page: Berggren et al. 2006 - Eocene Atlas, chap. 9, p. 310
References:
Berggren, W. A. (1977a). Atlas of Palaeogene Planktonic Foraminifera: some Species of the Genera Subbotina, Planorotalites, Morozovella, Acarinina and Truncorotaloides. In, Ramsay, A. T. S. (ed.) Oceanic Micropaleontology. Academic Press, London 205-300. gs
Berggren, W. A., Pearson, P. N., Huber, B. T. & Wade, B. S. (2006b). Taxonomy, biostratigraphy, and phylogeny of Eocene Acarinina. In, Pearson, P. N., Olsson, R. K., Hemleben, C., Huber, B. T. & Berggren, W. A. (eds) Atlas of Eocene Planktonic Foraminifera. Cushman Foundation for Foraminiferal Research, Special Publication . 41(Chap 9): 257-326. gsO
Blow, W. H. (1979). The Cainozoic Globigerinida: A study of the morphology, taxonomy, evolutionary relationships and stratigraphical distribution of some Globigerinida (mainly Globigerinacea). E. J. Brill, Leiden. 2: 1-1413. gs
Bolli, H. M. (1957d). The genera Globigerina and Globorotalia in the Paleocene-Lower Eocene Lizard Springs Formation of Trinidad. In, Loeblich, A. R. , Jr., Tappan, H., Beckmann, J. P., Bolli, H. M., Montanaro Gallitelli, E. & Troelsen, J. C. (eds) Studies in Foraminifera. U.S. National Museum Bulletin . 215: 61-82. gs
Fleisher, R. L. (1974a). Cenozoic planktonic foraminifera and biostratigraphy, Arabian Sea, Deep Sea Drilling Project, Leg 23A. Initial Reports of the Deep Sea Drilling Project. 23: 1001-1072. gsO
Hillebrandt, A. , von (1962). Das Paleozän und seine Foraminiferenfauna im Becken von Reichenhall und Salzburg. Abhandlungen Bayerischen Akademie der Wissenschaften. 108: 1-182. gs
Snyder, S. W. & Waters, V. J. (1985). Cenozoic planktonic foraminiferal biostratigraphy of the Goban Spur Region, Deep Sea Drilling Project Leg 80. Initial Reports of the Deep Sea Drilling Project. 80: 439-472. gs
Stainforth, R. M., Lamb, J. L., Luterbacher, H., Beard, J. H. & Jeffords, R. M. (1975). Cenozoic planktonic foraminiferal zonation and characteristics of index forms. University of Kansas Paleontological Contributions, Articles. 62: 1-425. gsO
Warraich, M. Y. & Ogasawara, K. (2001). Tethyan Paleocene-Eocene planktic foraminifera from the Rakhi Nala and Zinda Pir land sections of the Sulaiman Range, Pakistan. Science Reports of the Institute of Geosciences, University of Tsukuba. 22: 1-59. gs
Warraich, M. Y., Ogasawara, K. & Nishi, H. (2000). Late Paleocene to early Eocene planktic foraminiferal blostratigraphy of the Dungan Formation, Sulaiman Range, central Pakistan. Paleontological Research, Tokyo. 4(4): 275-301, 218 figures, 273 aendices. gs
Acarinina quetra compiled by the pforams@mikrotax project teamviewed: 25-4-2025