Globorotaliaapanthesma Loeblich and Tappan 1957a: 187, pl. 48: fig. la-c [Zone P4, Salt Mountain Limestone, Clarke Co., Alabama], pl. 55: fig. la-c [Zone P4, Vincentown Fm., New Jersey], pl. 58: fig. 4a-c (paratype), pl. 59: fig. la-c (holotype) [Zone P4, Aquia Fm., Virginia].
Globorotalia (Morozovella) apanthesma Loeblich and Tappan.—Jenkins, 1971:102, pl. 8: figs. 186-188 [Globigerinatriloculinoides Zone, Waipawan Stage, Middle Waipara River section, New Zealand].—Belford, 1984:9, pl. 16: figs. 1-8 [Zone P4, Papua, New Guinea].
? Globorotalia (Morozovella) apanthesma Loeblich and Tappan.—Blow,
1979:988, pl. 251: fig. 2 [Zone P7 of Blow, 1979 = Zone P6 this paper; Sample HK 1831, type level of G. rex Zone, "Carmen Jones Ravine," tributary of Cascas River, southern Trinidad].
Acarininaapanthesma (Loeblich and Tappan).—Huber, 1991b:446, pl. 4: figs. 1, 2 [Zone AP4, ODP Hole 738C/16R: 332.15 mbsf; Kerguelen Plateau, southern Indian Ocean].
Not Morozovellaapenthesma [sic] Lu and Keller, 1995:102, pl. 1: figs. 18, 19 [Zone P3b/P4x of Lu and Keller; DSDP Site 577/11/2: 52-54 cm; Shatsky Rise, northwestern Pacific Ocean]. [Olsson et al. 1999]
Taxonomic discussion: Blow (1979) drew attention to an important distinction within this plexus: apanthesma exhibits the essentially quadrate chamber pattern of the related (and descendant) aequa in the early whorls, whereas angulata (and related forms) retains the more "vorticiform," strongly recurved, early whorl pattern. In view of these, and other pertinent observations based on personal observations of type material (Blow, 1979), it is strange that he presented, almost as an afterthought, only a single, spiral view of a form assigned to apanthesma from (his) Zone P7 (= Zone P6 this paper) whose affinities are somewhat hard to evaluate. Earlier this species was considered a junior synonym of M. conicotruncata (Subbotina) by Luterbacher (1964) and of M. angulata (White) by Berggren (1977). Illustrated on Plate 11: Figures 10-12 is a specimen from the collections of Shutskaya (no. 645) in VNIGRI (St. Petersburg) that is probably referable to M. apanthesma. Although the slide containing this specimen is labeled as Globorotaliaangulata var. kubanensis Shutskaya, the illustration of the holotype resembles M. conicotruncata (Subbotina). Because the holotype in Moscow is lost, the identity of this taxon cannot be determined. Further confusing the taxonomic status of Shutskaya's taxon is the other specimen from the same slide (no. 645) illustrated on Plate 11: Figures 13-15. This specimen is probably referable to M. acutispira. [Olsson et al. 1999]
Distinguishing features: Parent taxon (Morozovella): Test typically plano-convex, chambers strongly anguloconical. Wall strongly pustulose (muricate) on parts of spire and umbilicus. Most species with muricocarina. This taxon: Planoconvex, umbilicoconvex, test with lobulate, weakly muricocarinate periphery; 4-5 chambers in last whorl, inflated to subangular on umbilical side, moderately convex, triangular (lunate) in edge view; intercameral sutures on umbilical side depressed, radially curved and slightly depressed on spiral side; umbilical side distinctly muricate, coarsely perforate on spiral side; umbilicus relatively narrow, deep; aperture an interiomarginal, umbilical-extraumbilical arch with narrow, continuous intraperiumbilical lip.
NB These concise distinguishing features statements are used in the tables of daughter-taxa to act as quick summaries of the differences between e.g. species of one genus. They are being edited as the site is developed and comments on them are especially welcome.
Description
Diagnostic characters: Planoconvex, umbilicoconvex, test with lobulate, weakly muricocarinate periphery; 4-5 chambers in last whorl, inflated to subangular on umbilical side, moderately convex, triangular (lunate) in edge view; intercameral sutures on umbilical side depressed, radially curved and slightly depressed on spiral side; umbilical side distinctly muricate, coarsely perforate on spiral side; umbilicus relatively narrow, deep; aperture an interiomarginal, umbilical-extraumbilical arch with narrow, continuous intraperiumbilical lip. [Olsson et al. 1999]
Character matrix
test outline:
Lobate
chamber arrangement:
Trochospiral
edge view:
Planoconvex
aperture:
Umbilical-extraumbilical
sp chamber shape:
Crescentic
coiling axis:
High
periphery:
Muricocarinate
aperture border:
Thin lip
umb chbr shape:
Subtriangular
umbilicus:
Wide
periph margin shape:
Subangular
accessory apertures:
None
spiral sutures:
Weakly depressed
umb depth:
Deep
wall texture:
Spinose
shell porosity:
Macroperforate: >2.5µm
umbilical or test sutures:
Strongly depressed
final-whorl chambers:
4-5
N.B. These characters are used for advanced search. N/A - not applicable
Biogeography and Palaeobiology
Geographic distributionNorthern middle latitudes to the Southern Ocean. [Olsson et al. 1999]
Aze et al. 2011 summary: Northern middle latitudes to the Southern Ocean; based on Olsson et al. (1999) Isotope paleobiologyLimited isotopic data suggest M. apanthesma is similar in δ18O and δ13C to other morozovellids and has a more positive δ13C and more negative δ18O than coexisting Globanomalina and Subbotina (Lu and Keller, 1996; Berggren and Norris, 1997). [Olsson et al. 1999] Aze et al. 2011 ecogroup 1 - Open ocean mixed-layer tropical/subtropical, with symbionts. Based on very heavy _13C and relatively light _18O. Sources cited by Aze et al. 2011 (appendix S3): Lu & Keller (1996); Berggren & Norris (1997) Phylogenetic relationsMorozovellaapanthesma and its descendants, M. aequa and M. subbotinae, differ from the M. angulata-M. velascoensis group in possessing a relatively even distribution of fine muricae (pustules) over the surface of the test in contrast to the development of muricae-free surfaces between the muricorcarina and adumbilical ridges in the M. velascoensis plexus. Morozovellaapanthesma shares characteristics intermediate between M. praeangulata (its antecedent) and M. aequa (its descendant). The former is generally somewhat smaller (contra Loeblich and Tappan, 1957a; see also Jenkins, 1971), with more subdued muricate wall texture and more strongly recurved chambers when seen from the spiral surface. Compared to its ancestor, M. aequa exhibits a reduced number of chambers and a more anguloconical test. [Olsson et al. 1999]
Geological Range: Notes: Zone P3b to Zone P4c. [Olsson et al. 1999] Last occurrence (top): at top of P4c subzone (100% up, 57.1Ma, in Thanetian stage). Data source: Olsson et al. (1999) f5a First occurrence (base): at base of P3b subzone (0% up, 61.3Ma, in Selandian stage). Data source: Olsson et al. (1999) f5a
Plot of occurrence data:
Range-bar - range as quoted above, pink interval top occurs in, green interval base occurs in.
Triangles indicate an event for which a precise placement has been suggested
Histogram - Neptune occurrence data from DSDP and ODP proceedings. Pale shading <50 samples in time bin. Interpret with caution & read these notes
Primary source for this page: Olsson et al. 1999 - Atlas of Paleocene Planktonic Foraminifera, p. 59
References:
Berggren, W. A. (1977a). Atlas of Palaeogene Planktonic Foraminifera: some Species of the Genera Subbotina, Planorotalites, Morozovella, Acarinina and Truncorotaloides. In, Ramsay, A. T. S. (ed.) Oceanic Micropaleontology. Academic Press, London 205-300. gs
Blow, W. H. (1979). The Cainozoic Globigerinida: A study of the morphology, taxonomy, evolutionary relationships and stratigraphical distribution of some Globigerinida (mainly Globigerinacea). E. J. Brill, Leiden. 2: 1-1413. gs
Loeblich, A. R. & Tappan, H. (1957b). Planktonic foraminifera of Paleocene and early Eocene Age from the Gulf and Atlantic coastal plains. In, Loeblich, A. R. , Jr., Tappan, H., Beckmann, J. P., Bolli, H. M., Montanaro Gallitelli, E. & Troelsen, J. C. (eds) Studies in Foraminifera. U.S. National Museum Bulletin . 215: 173-198. gs
Luterbacher, H. P. (1964). Studies in some Globorotalia from the Paleocene and Lower Eocene of the Central Apennines. Eclogae Geologicae Helvetiae. 57: 631-730. gsO
Olsson, R. K., Hemleben, C., Berggren, W. A. & Huber, B. T. (1999). Atlas of Paleocene Planktonic Foraminifera. Smithsonian Institution Press, Washington, DC. (85): 1-252. gs
Ouda, K. & Aubry, M. -P. (2003). The Upper Paleocene- Lower Eocene of the Upper Nile Valley: Part 1: Stratigraphy. Micropaleontology. 49(Supplement 1): 212-. gs
Morozovella apanthesma compiled by the pforams@mikrotax project teamviewed: 15-11-2025