Globorotaliacrassata (Cushman).—Glaessner,1937:31, pl. 1: figs. 7a-c [lower Eocene Zone of compressed globorotaliids, Subzone with Globorotaliamarginodentata, Foraminiferal Beds, Series F1, near Nal’chik, northern Caucasus, former Soviet Union].— Subbotina, 1947:119-121 (partim), not pl. 5: figs. 7-12 = ?M. subbotinae (Morozova), pl. 9: figs. 15-17 [Globorotalia ex gr. canariensis, later (1953) designated the Zone of compressed globorotaliids, Globorotaliamarginodentata Subzone, Kutaiss Horizon (F2), Kadiizhensk Region, Psish River, northern Caucasus, former Soviet Union]. [Not Cushman, 1925.]
Globorotaliamarginodentata Subbotina, 1953:212, pl. 17: figs. 14 [lower Eocene, Zone of compressed globorotaliids, Subzone with Globorotaliamarginodentata, Foraminiferal Beds, Series F1, near Nal’chik, northern Caucasus, former Soviet Union [refigured from Glaessner, 1937:31, pl. 7: figs. 7a-c; pl. 17, figs. 15a-c (holotype), and 16a-c, lower Eocene Zone of compressed globorotaliids, Subzone with Globorotaliamarginodentata, base of Foraminiferal Beds, Green Series, Kuban River section, northern Caucasus, former Soviet Union]; pl. 18: figs. 1a-c [lower Eocene Zone of compressed globorotaliids, Subzone with Globorotaliamarginodentata, Foraminiferal Beds, base of Kutais Horizon (F2), Khadyzhensk District, Psish River section, northern Caucasus, former Soviet Union]; pl. 18: figs. 2a-c [lower Eocene Zone of conical globorotaliids, Foraminiferal Beds, Kutai Horizon (F2), Gubs River section, Barakaevsk District, northern Caucasus, former Soviet Union]; pl. 18: figs. 3a-c [lower Eocene Zone of compressed globorotaliids, Foraminiferal Beds, Series F1, near Nal’chik, Khieu River section, northern Caucasus; interpreted as transitional form to G. lensiformis].— Luterbacher, 1964:673, text-figs. 75a-76c [lower Eocene Zone of compressed globorotaliids, Khieu River section, northern Caucasus, former Soviet Union, determined/ identified by N .N. Subbotina, fide Luterbacher, 1964:674]; 77a-78c [uppermost Paleocene Globorotaliavelascoensis Zone, Velasco Fm., Ebano, Mexico; referred to as “aff. marginodentata”]; 79a-c [lower Eocene G. velascoensis Zone, Gubbio section, northern Apennines, Italy; referred to as “aff. marginodentata”]; 80a-c [lower Eocene G. velascoensis Zone, Gubbio section, Italy]; 81a-82c [lower Eocene G. aequa Zone, Gubbio section, Italy]; 83a-c [lower Eocene G. aequa Zone, Gubbio section, Italy; referred to as “intermediate between Globorotaliamarginodentata and Globorotaliasubbotinae”]; 84a-c [lower Eocene G. formosaformosa /G. subbotinae Zone, Gubbio section, Italy]. —Samanta, 1970:626, pl. 96: figs. 3,4 [lower Eocene, Pondicherry Fm., Pondicherry, south east India].—Luterbacher, 1975a:727, pl. 2: figs. 6a-c [lower Eocene Zone P6, DSDP Hole 313, Mid-Pacific Mountains].—Luterbacher, 1975b:65, pl. 4: figs. 4-6 [lower Eocene G. subbotinae Zone, Possagno section, Italy].
Globorotalia (Truncorotalia) aequamarginodentata Subbotina.—Hillebrandt, 1962:135, pl. 13: figs. 9a-11 [lower Eocene, Zone G, Reichenhall-Salzburg Basin, Germany].
Truncorotaliamarginodentatamarginodentata (Subbotina).—Gohrbandt, 1963:62, pl. 6: figs. 4-6 [lower Eocene, Zone F, near Salzburg, Austria].
Globorotalia (Morozovella) aequamarginodentata Subbotina. —Jenkins, 1971:101, text-figs. 177-179 [uppermost Paleocene G. (S.) triloculinoides Zone, Waipawan Stage, Middle Waipara River section, North Island, NZ].
Morozovellamarginodentata (Subbotina).—Berggren, 1971:76, pl. 5: fig. 9 [lower Eocene Zone P7, DSDP Hole 20C, South Atlantic Ocean].—Berggren, 1977:241, chart No. 8 [specimens refigured from the literature].—Snyder and Waters, 1985:460, pl. 8, figs. 13a-14c [lower Eocene Zone P7, DSDP Hole 549, Goban Spur, Atlantic Ocean].— Warraich, Ogasawara and Nishi, 2000:293, figs. 17. 4,5,10 [lower Eocene Zone P7, Dungan Fm., Rakhi Nala River, Sulaiman Range, Pakistan].—Warraich and Ogasawara, 2001:40, fig. 10. 7-9 [lower Eocene Zone P7, Dungan Fm., Rakhi Nala River, Sulaiman Range, Pakistan].
Globorotalia (Morozovella) subbotinae (s.l) forma marginodentata Subbotina.—Blow, 1979:1024-1026, pl. 139: figs. 1-9 and pl. 140: figs.1-3 [lower Eocene Zone P8b, DSDP Hole 20C, South Atlantic Ocean; see also pl. 223: figs. 5, 6 and pl. 224: figs. 1 and 2].
Truncorotaliamarginodentataaperta Gohrbandt, 1963:62, pl. 5: figs. 10-15 [Zones E and F, near Salzburg, Austria].
Taxonomic discussion: Placement of this taxon is extremely difficult. “Typical” specimens of marginodentata are denoted by their marginally compressed/scalloped chambers and thickly muricate periphery. However, in most early Eocene (sub)tropical fossil assemblages intergradation of these morphologies with “typical” morphologies of M. gracilis and M. subbotinae exist. Blow (1979, p. 1025, 1026) drew attention to the development of marginodentate ornament on both a subbotinae and gracilis morphotype, interpreted the marginodentate forms as “extreme phenotypes developed under particular environmental conditions...” and observed that “the ‘marginodentate’ forms appear to occur only in areas of high biotic productivity”. He chose to view these ecophenotypes nomenclatorially as extreme forms of subbotinae (sensu stricto) and denoted them as G. (M.) subbotinae forma marginodentata. Berggren (1971, p. 76) expressed similar views in suggesting that marginodentata may be an ecophenotype of gracilis. We retain the name marginodentata here for forms possessing the distinctly thick and broad peripheral muricocarina. [Berggren & Pearson 2006]
Distinguishing features: Parent taxon (Morozovella): Test typically plano-convex, chambers strongly anguloconical. Wall strongly pustulose (muricate) on parts of spire and umbilicus. Most species with muricocarina. This taxon: Morphotype, with pronounced peripheral compression of the chambers on the umbilical side and a thick muricocarina formed form long muricae.
NB These concise distinguishing features statements are used in the tables of daughter-taxa to act as quick summaries of the differences between e.g. species of one genus. They are being edited as the site is developed and comments on them are especially welcome.
Description
Morphology: Low trochospiral, biconvex, oval to subcircular test with lobulate, thick and strongly fimbriate muricocarina; chambers on umbilical side moderately inflated, depressed towards periphery; flat on spiral side; 4.5 - 5 chambers in last whorl, (sub)triangular, compressed along peripheral margin, rising gradually towards umbilicus; sutures straight to weakly recurved, radial, depressed; aperture a low, umbilical-extraumbilical arch or slit extending towards, but not to, the periphery; umbilicus narrow, deep; in spiral view approximately 10-12 chambers in 2.5 whorls; early chambers heavily muricate and elevated above test surface; chambers lens-shaped to semicircular, intercameral sutures curved, strongly muricate, elevated above test surface; scalloped peripheral muricocarinate periphery characterized by dense concentration of conical or long slender muricae and the fusion/coalescence of muricae; in edge view low to moderately biconvex; early chambers elevated above later whorl(s); low to medium conical angle (< 45° to ~75°). [Berggren & Pearson 2006] Wall type: Densely muricate on both sides, nonspinose, normal perforate. [Berggren & Pearson 2006] Size: Holotype Diameter: 0.43 mm; thickness/width: 0.24 mm (Subbotina, 1953, p. 213) [Berggren & Pearson 2006]
Character matrix
test outline:
Subcircular
chamber arrangement:
Trochospiral
edge view:
Equally biconvex
aperture:
Umbilical-extraumbilical
sp chamber shape:
Petaloid
coiling axis:
Low
periphery:
Muricocarinate
aperture border:
N/A
umb chbr shape:
Subtriangular
umbilicus:
Narrow
periph margin shape:
Subangular
accessory apertures:
None
spiral sutures:
Raised muricate
umb depth:
Deep
wall texture:
Coarsely muricate
shell porosity:
Finely Perforate: 1-2.5µm
umbilical or test sutures:
Moderately depressed
final-whorl chambers:
4.5-5
N.B. These characters are used for advanced search. N/A - not applicable
Biogeography and Palaeobiology
Geographic distributionWidely distributed in (sub)tropical regions (Caribbean, Atlantic Ocean, Indo-Pacific region, North Caucasus) [Berggren & Pearson 2006]
Aze et al. 2011 summary: Low latitudes; based on Berggren & Pearson (2006) Isotope paleobiologyNo data available. [Berggren & Pearson 2006] Aze et al. 2011 ecogroup 1 - Open ocean mixed-layer tropical/subtropical, with symbionts; based on comparison with other species of the genus. Phylogenetic relationsThis morphotype is closely related to the subbotinae-gracilis group. It probably evolved from M. subbotinae and does not appear to have left any descendants. [Berggren & Pearson 2006]
Most likely ancestor:Morozovella subbotinae - at confidence level 3 (out of 5). Data source: Berggren & Pearson (2006) f11.1.
Biostratigraphic distribution
Geological Range: Notes: Zone P5 to Zone E5. [Berggren & Pearson 2006] Last occurrence (top): in mid part of E5 zone (50% up, 51.6Ma, in Ypresian stage). Data source: Berggren & Pearson (2006) f11.1 First occurrence (base): in upper part of P5 zone (80% up, 56.2Ma, in Thanetian stage). Data source: Berggren & Pearson (2006) f11.1
Plot of occurrence data:
Range-bar - range as quoted above, pink interval top occurs in, green interval base occurs in.
Triangles indicate an event for which a precise placement has been suggested
Histogram - Neptune occurrence data from DSDP and ODP proceedings. Pale shading <50 samples in time bin. Interpret with caution & read these notes
Primary source for this page: Berggren & Pearson 2006 - Eocene Atlas, chap. 11, p. 368
References:
Berggren, W. A. & Pearson, P. N. (2006a). Taxonomy, biostratigraphy, and phylogeny of Eocene Morozovella. In, Pearson, P. N., Olsson, R. K., Hemleben, C., Huber, B. T. & Berggren, W. A. (eds) Atlas of Eocene Planktonic Foraminifera. Cushman Foundation for Foraminiferal Research, Special Publication . 41(Chap 11): 343-376. gsO
Berggren, W. A. (1971c). Paleogene planktonic foraminiferal faunas on Legs I-IV (Atlantic Ocean) JOIDES Deep Sea Drilling Program: a synthesis. In, Farinacci, A. (ed.) Proceedings of the Second Planktonic Conference, Roma 1970. Edizioni Tecnoscienza, Rome 57-77. gs
Berggren, W. A. (1977a). Atlas of Palaeogene Planktonic Foraminifera: some Species of the Genera Subbotina, Planorotalites, Morozovella, Acarinina and Truncorotaloides. In, Ramsay, A. T. S. (ed.) Oceanic Micropaleontology. Academic Press, London 205-300. gs
Blow, W. H. (1979). The Cainozoic Globigerinida: A study of the morphology, taxonomy, evolutionary relationships and stratigraphical distribution of some Globigerinida (mainly Globigerinacea). E. J. Brill, Leiden. 2: 1-1413. gs
Cushman, J. A. (1925e). Some new foraminifera from the Velasco shale of Mexico. Contributions from the Cushman Laboratory for Foraminiferal Research. 1(1): 18-23. gs
Glaessner, M. F. (1937a). Planktonforaminiferen aus der Kreide und dem Eozän und ihre stratigraphische Bedeutung. Etyudy po Mikropaleontologiy, Paleontologicheskaya Laboratoriya Moskovskogo Gosudarstvennogo Universiteta. 1(1): 27-46. gs
Gohrbandt, K. (1963). Zur Gliederung des Palaeogen im Helvetikum nordlich Salzburg nach planktonischen Foraminiferen. Mitteilungen der Geologischen Gesellschaft in Wien. 56(1): 63-. gs
Hillebrandt, A. , von (1962). Das Paleozän und seine Foraminiferenfauna im Becken von Reichenhall und Salzburg. Abhandlungen Bayerischen Akademie der Wissenschaften. 108: 1-182. gs
Jenkins, D. G. (1971). New Zealand Cenozoic Planktonic Foraminifera. New Zealand Geological Survey, Paleontological Bulletin. 42: 1-278. gs
Luterbacher, H. P. (1964). Studies in some Globorotalia from the Paleocene and Lower Eocene of the Central Apennines. Eclogae Geologicae Helvetiae. 57: 631-730. gsO
Luterbacher, H. P. (1975a). Paleocene and Early Eocene planktonic foraminifera Leg 32, Deep Sea Drilling Project. Initial Reports of the Deep Sea Drilling Project. 32: 725-728. gs
Luterbacher, H. P. (1975b). Planktonic Foraminifera of the Paleocene and Early Eocene, Possagno Section. Schweizerische Paläontologische Abhandlungen. 97: 57-67. gs
Snyder, S. W. & Waters, V. J. (1985). Cenozoic planktonic foraminiferal biostratigraphy of the Goban Spur Region, Deep Sea Drilling Project Leg 80. Initial Reports of the Deep Sea Drilling Project. 80: 439-472. gs
Subbotina, N. N. (1953). Foraminiferes fossiles d'URSS Globigerinidae, Globorotaliidae, Hantkeninidae. Bureau de Recherches Geologiques et Minieres. 2239: 1-144. gs
Warraich, M. Y. & Ogasawara, K. (2001). Tethyan Paleocene-Eocene planktic foraminifera from the Rakhi Nala and Zinda Pir land sections of the Sulaiman Range, Pakistan. Science Reports of the Institute of Geosciences, University of Tsukuba. 22: 1-59. gs
Warraich, M. Y., Ogasawara, K. & Nishi, H. (2000). Late Paleocene to early Eocene planktic foraminiferal blostratigraphy of the Dungan Formation, Sulaiman Range, central Pakistan. Paleontological Research, Tokyo. 4(4): 275-301, 218 figures, 273 aendices. gs
Morozovella marginodentata compiled by the pforams@mikrotax project teamviewed: 14-1-2025
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holotype: G. marginodentata
