pforams@mikrotax - Morozovella velascoensis pforams@mikrotax - Morozovella velascoensis

Morozovella velascoensis

Classification: pf_cenozoic -> Truncorotaloididae -> Morozovella -> Morozovella velascoensis
Sister taxa: M. caucasica, M. crater, M. aragonensis, M. lensiformis ⟩⟨ M. marginodentata, M. formosa, M. gracilis, M. subbotinae, M. aequa, M. apanthesma ⟩⟨ M. edgari, M. allisonensis, M. acuta, M. occlusa, M. acutispira, M. pasionensis, M. velascoensis, M. conicotruncata, M. angulata, M. praeangulata, M. sp.


Citation: Morozovella velascoensis (Cushman 1925)
taxonomic rank: Species
Basionym: Pulvinulina velascoensis
Taxonomic discussion: This is the type species of the genus Morozovella McGowran (in Luterbacher, 1964) and one of the most characteristic and easily recognized Paleocene morozovellids. It has been assigned to various genera and subgenera over the past half century but would appear to have found a suitable home in the genus Morozovella as defined by McGowran in Luterbacher (1964; see also McGowran, 1968).
This species has been confused with the heterochronously isomorphic form M. caucasica (Glaessner) (see Subbotina, 1953, pl. 19: figs. la-2c, who identified strongly muricocarinate and circumumbilically muricate forms from the lower Eocene zone of conical globorotaliids of the northern Caucasus as G. velascoensis; see also El-Naggar, 1966, who was equally confused by the distinctions between these two taxa, and Blow, 1979, for clarification of the problems associated with separation and recognition of these two forms). [Olsson et al. 1999]

Catalog entries: Pulvinulina velascoensis

Type images:

Distinguishing features:
Parent taxon (Morozovella): Test typically plano-convex, chambers strongly anguloconical.
Wall strongly pustulose (muricate) on parts of spire and umbilicus. Most species with muricocarina.

This taxon: Like M. conicotruncata but with coarsely muricate adumbilical ridges and muricocarina; and complete loss of muricae on the chamber surfaces.

NB These concise distinguishing features statements are used in the tables of daughter-taxa to act as quick summaries of the differences between e.g. species of one genus.
They are being edited as the site is developed and comments on them are especially welcome.


Character matrix
test outline:Subcircularchamber arrangement:Trochospiraledge view:Planoconvexaperture:Umbilical-extraumbilical
sp chamber shape:Crescenticcoiling axis:Highperiphery:Muricocarinateaperture border:Thin flange
umb chbr shape:Subtriangularumbilicus:Narrowperiph margin shape:Subangularaccessory apertures:None
spiral sutures:Raised muricateumb depth:Deepwall texture:Moderately muricateshell porosity:Finely Perforate: 1-2.5µm
umbilical or test sutures:Moderately depressedfinal-whorl chambers:6.5-7.5 N.B. These characters are used for advanced search. N/A - not applicable

Biogeography and Palaeobiology

Geographic distribution

Morozovella velascoensis has a wide geographic distribution but is a predominantly (sub)tropical to temperate form; it has not been recorded from high northern or southern (subantarctic) latitudes (> 45° N or S). The disappearance of this taxon is a distinct biostratigraphic event that is used to define the boundary between Zones P5 and
P6, which occurs in mid-Chron C24r, and is closely correlative with the Paleocene/Eocene boundary as denoted in the Belgian and/or London-Hampshire Basin(s) of northwestern Europe (Berggren and Aubry, 1996) (Figure 25). [Olsson et al. 1999]
Aze et al. 2011 summary: Low to middle latitudes; based on Olsson et al. (1999)

Isotope paleobiology
Morozovella velascoensis has δ13C similar to coexisting species of Morozovella and Acarinina and more positive δ13C and more negative δ18O than Subbotina (Boersma and Premoli Silva, 1983; Shackleton et al., 1985; D'Hondt et al., 1994). Morozovella velascoensis displays a pronounced increase in 8UC with increased test size but little corresponding change in δ18O (D'Hondt et al., 1994). This species typically has more negative δ18O than all other analyzed species of Morozovella (Berggren and Norris, 1997). [Olsson et al. 1999]
Aze et al. 2011 ecogroup 1 - Open ocean mixed-layer tropical/subtropical, with symbionts. Based on very heavy _13C and relatively light _18O. Sources cited by Aze et al. 2011 (appendix S3): Berggren & Norris (1997)

Phylogenetic relations
This species evolved from Morozovella conicotruncata (Subbotina) within Subzone P3b by the acquisition of coarsely muricate adumbilical ridges, the complete loss of muricae on the spiral chamber surfaces and between the muricocarina and umbilical collar, as well as the formation of a thick, coarsely muricate muricocarina. [Olsson et al. 1999]

Most likely ancestor: Morozovella conicotruncata - at confidence level 4 (out of 5). Data source: Olsson et al. (1999) f5a.
Likely descendants: Morozovella acuta; Morozovella allisonensis; Morozovella edgari; Morozovella pasionensis; plot with descendants

Biostratigraphic distribution

Geological Range:
Notes: Zone P3b to Zone P5 (top). [Olsson et al. 1999]
The LAD of Morozovella velascoensis marks the base of zone E3 / top of E2 (Wade et al. 2011)
Last occurrence (top): at top of E2 zone (100% up, 55.2Ma, in Ypresian stage). Data source: zonal marker (Wade et al. 2011)
First occurrence (base): near base of P3b subzone (10% up, 61.3Ma, in Selandian stage). Data source: Olsson et al. 1999

Plot of occurrence data:

Primary source for this page: Olsson et al. 1999 - Atlas of Paleocene Planktonic Foraminifera, p. 66


Berggren, W. A. & Pearson, P. N. (2006a). Taxonomy, biostratigraphy, and phylogeny of Eocene Morozovella. In, Pearson, P. N., Olsson, R. K., Hemleben, C., Huber, B. T. & Berggren, W. A. (eds) Atlas of Eocene Planktonic Foraminifera. Cushman Foundation for Foraminiferal Research, Special Publication . 41(Chap 11): 343-376. gs O

Blow, W. H. (1979). The Cainozoic Globigerinida: A study of the morphology, taxonomy, evolutionary relationships and stratigraphical distribution of some Globigerinida (mainly Globigerinacea). E. J. Brill, Leiden. 2: 1-1413. gs

Cushman, J. A. (1925e). Some new foraminifera from the Velasco shale of Mexico. Contributions from the Cushman Laboratory for Foraminiferal Research. 1(1): 18-23. gs

El-Naggar, Z. R. (1966). Stratigraphy and planktonic foraminifera of the Upper Cretaceous-Lower Tertiary succession in the Esna-Idfu region, Nile Valley, Egypt, U. A. R. Bulletin of the British Museum (Natural History). supplement 2: 1-291. gs

Luterbacher, H. P. (1964). Studies in some Globorotalia from the Paleocene and Lower Eocene of the Central Apennines. Eclogae Geologicae Helvetiae. 57: 631-730. gs O

McGowran, B. J. (1968). Reclassification of Early Tertiary Globorotalia. Micropaleontology. 14: 179-198. gs

Olsson, R. K., Hemleben, C., Berggren, W. A. & Huber, B. T. (1999). Atlas of Paleocene Planktonic Foraminifera. Smithsonian Institution Press, Washington, DC. 1-252. gs

Postuma, J. A. (1971). Manual of planktonic foraminifera. Elsevier for Shell Group, The Hague. 1-406. gs

Subbotina, N. N. (1953). Foraminiferes fossiles d'URSS Globigerinidae, Globorotaliidae, Hantkeninidae. Bureau de Recherches Geologiques et Minieres. 2239: 1-144. gs

Wade, B. S., Pearson, P. N., Berggren, W. A. & Pälike, H. (2011). Review and revision of Cenozoic tropical planktonic foraminiferal biostratigraphy and calibration to the geomagnetic polarity and astronomical time scale. Earth-Science Reviews. 104: 111-142. gs


Morozovella velascoensis compiled by the pforams@mikrotax project team viewed: 23-7-2024

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Comments (2)

Le Coze


In the Biostratigraphic distribution, Taxon plotted, Subbotina velascoensis is mentionned but it is a different species (= Globigerina velascoensis Cushman, 1925 not Pulvinulina velascoensis Cushman, 1925).



Jeremy Young(UK)

Hi Francois - thank you very much, and well spotted.