Globorotaliacrassata (Cushman).—Cushman and Barksdale, 1930:67-68, pl. 12: fig. 7a-b [Eocene, Martinez Fm., California].—Cushman, 1939:74, pl. 12: fig. 19 [“upper Eocene”, submarine core off eastern North America]. [Not Cushman, 1925.]
Morozovellacrassata (Cushman).—Pearson, Shackleton and Hall, 1993, pl. 2: figs. 4-6 [middle Eocene Zone P11-P12, DSDP Site 523, Walvis Ridge, south Atlantic Ocean]. [Not Cushman, 1925.]
Morozovellabandyi Fleisher, 1974:1034, pl. 14: figs. 3-8 [Middle Zone P10 to Zone P11, DSDP Sites 219 and 220, Arabian Sea].
Taxonomic discussion: Morozovelloidesbandyi is a common form in Zones E8 and E9, but first appears in the upper part of Zone E7 where it intergrades with Acarininapraetopilensis. It has probably been described previously on many occasions as crassata or spinulosa, but such synonymies in the older literature are difficult to verify because M. bandyi is strongly homeomorphic to the Morozovellaaequa – M. subbotinae group of the lower Eocene. [Pearson & Berggren 2006]
Distinguishing features: Parent taxon (Morozovelloides): Like Morozovella, but derived from Acarinina. This taxon: Distinguished from all other species of Morozovelloides by the strongly disjunct long axes of the chambers, as seen in spiral view.
NB These concise distinguishing features statements are used in the tables of daughter-taxa to act as quick summaries of the differences between e.g. species of one genus. They are being edited as the site is developed and comments on them are especially welcome.
Description
Diagnostic characters: Distinguished from all other species of Morozovelloides by the strongly disjunct long axes of the chambers, as seen in spiral view, and from Acarininapraetopilensis by having a more dorso-ventrally flattened test and peripheral (as opposed to circum-cameral) muricocarina. [Pearson & Berggren 2006] Morphology: Planoconvex to lenticular test, coiled in a low trochospiral, with concentration of bladed muricae around the petaloid periphery that constitute a discontinuous muricocarina. Four to five chambers in the final whorl, added in such a way that their long axes are strongly disjunct when viewed on the dorsal side. Typically the chambers are imbricate, sometimes strongly so, as if tilted relative to the dorso-ventral plane. Small supplementary apertures are commonly present at the base of the sutures between the penultimate and antepenultimate chambers, and more rarely earlier in the spiral. Primary aperture is a low, flat arch, in an umbilical-extraumbilical position. Umbilicus moderately broad and deep. Ventral sutures depressed with umbilical shoulders of chambers commonly showing concentrations of muricae (but not to the extent seen in M. coronata, see below). [Pearson & Berggren 2006] Wall type: Muricate, normal perforate with smooth areas on the spiral side. [Pearson & Berggren 2006] Size: Maximum diameter of holotype about 0.28 mm; height 0.16 mm. [Pearson & Berggren 2006]
Character matrix
test outline:
Lobate
chamber arrangement:
Trochospiral
edge view:
Planoconvex
aperture:
Umbilical-extraumbilical
sp chamber shape:
Crescentic
coiling axis:
Low
periphery:
Muricocarinate
aperture border:
N/A
umb chbr shape:
Subtriangular
umbilicus:
Wide
periph margin shape:
Subangular
accessory apertures:
None
spiral sutures:
Raised muricate
umb depth:
Deep
wall texture:
Moderately muricate
shell porosity:
Finely Perforate: 1-2.5µm
umbilical or test sutures:
Moderately depressed
final-whorl chambers:
4-5
N.B. These characters are used for advanced search. N/A - not applicable
Biogeography and Palaeobiology
Geographic distributionCosmopolitan. [Pearson & Berggren 2006]
Aze et al. 2011 summary: Cosmopolitan; based on Pearson & Berggren (2006) Isotope paleobiologyOxygen and carbon isotopes indicate a mixed layer, phososymbiotic habitat (Boersma and others, 1987 [recorded as Morozovella cf. aequa], Pearson and others, 1993 and Pearson and others, 2001 [recorded as Morozovellacrassata]). Boron isotope data suggest a deeper habitat than co-occurring M. crassatus (Pearson and Palmer, 1999, recorded as Morozovellacrassata; Morozovelloidescrassatus recorded as Morozovellaspinulosa). [Pearson & Berggren 2006] Aze et al. 2011 ecogroup 1 - Open ocean mixed-layer tropical/subtropical, with symbionts. Based on very heavy _13C and relatively light _18O. Sources cited by Aze et al. 2011 (appendix S3): Boersma et al. (1987); Pearson et al. (1993, 2001a) Phylogenetic relationsMorozovelloidesbandyi evolved from Acarininapraetopilensis and gave rise to M. crassatus. [Pearson & Berggren 2006]
Geological Range: Notes: First appears in the upper part of Zone E7, intergrading with A. praetopilensis. Most common in Zones E8 and E9, but ranging as high as Zone E10. [Pearson & Berggren 2006] NB Given the problems with zones E7 and E8 disccussed by Wade et al. 2100 this range may need revising [JRY 2017] Last occurrence (top): within E10 zone (41.89-43.23Ma, top in Lutetian stage). Data source: Pearson & Berggren 2006, f10.1 First occurrence (base): within E7b subzone (45.72-48.31Ma, base in Ypresian stage). Data source: Pearson & Berggren 2006, f10.1
Plot of occurrence data:
Range-bar - range as quoted above, pink interval top occurs in, green interval base occurs in.
Triangles indicate an event for which a precise placement has been suggested.
Grey shading between taxa indicates intergrading species within an anagenetic lineage (data from
Lamyman et al. 2026).
Histogram - Neptune occurrence data from DSDP and ODP proceedings. Pale shading <50 samples in time bin. Interpret with caution & read these notes
Taxon plotted: Morozovelloides bandyi, synonyms included - Morozovelloides bandyi;
Primary source for this page: Pearson & Berggren 2006 - Eocene Atlas, chap. 10, p. 330
References:
Beckmann, J. P. (1953). Die Foraminiferen der Oceanic Formation (Eocaen-Oligocaen) von Barbados, Kl. Antillen. Eclogae Geologicae Helvetiae. 46(2): 301-412. gsO
Boersma, A., Premoli Silva, I. & Shackleton, N. J. (1987). Atlantic Eocene planktonic foraminiferal paleohydrographic indicators and stable isotope paleoceanography. Paleoceanography. 2: 287-331. gs
Cushman, J. A. & Barksdale, J. D. (1930). Eocene foraminifera from Martinez, California. Contributions Department of Geology, Stanford University. 1: 55-73. gs
Cushman, J. A. (1925b). An Eocene fauna from the Moctezuma River, Mexico. Bulletin of the American Association of Petroleum Geologists. 9(2): 298-301. gs
Cushman, J. A. (1927b). New and interesting foraminifera from Mexico and Texas. Contributions from the Cushman Laboratory for Foraminiferal Research. 3(2): 111-119. gs
Cushman, J. A. (1939b). Eocene foraminifera from submarine cores off the eastern coast of North America. Contributions from the Cushman Laboratory for Foraminiferal Research. 15(3): 49-76. gs
Fleisher, R. L. (1974a). Cenozoic planktonic foraminifera and biostratigraphy, Arabian Sea, Deep Sea Drilling Project, Leg 23A. Initial Reports of the Deep Sea Drilling Project. 23: 1001-1072. gsO
Pearson, P. N. & Berggren, W. A. (2006). Taxonomy, biostratigraphy, and phylogeny of Morozovelloides n. gen. In, Pearson, P. N., Olsson, R. K., Hemleben, C., Huber, B. T. & Berggren, W. A. (eds) Atlas of Eocene Planktonic Foraminifera. Cushman Foundation for Foraminiferal Research, Special Publication . 41(Chap 10): 327-342. gsO
Pearson, P. N. & Palmer, M. R. (1999). Middle Eocene seawater pH and atmospheric carbon dioxide concentrations. Science. 284: 1824-1826. gs
Pearson, P. N., Shackleton, N. J. & Hall, M. A. (1993). Stable isotope paleoecology of middle Eocene planktonic foraminifera and multi-species isotope stratigraphy, DSDP Site 523, South Atlantic. Journal of Foraminiferal Research. 23: 123-140. gs
Morozovelloides bandyi compiled by the pforams@mikrotax project teamviewed: 9-10-2026