Hastigerinellacolombiana Petters, 1954:40, pl. 80: figs. 10a-b [middle Eocene, Asterigerinacrassaformis zone, upper Carreto Fm., Colombia].—Weiss, 1955:309, pl. 2: figs. 9-10 [middle Eocene, Talara, Chira, and Mirador Fms, Peru].
Clavigerinella alicantensis Cremades Campos 1979 [according to P.N. Pearson - email comm. 5 May 2023]
Taxonomic discussion: Complete specimens are very rare; the holotype illustrated by Petters, and shown here in SEM for the first time, is one of very few that has been isolated more or less intact. The species is usually identified by the isolated, paddle-shaped chambers. A few complete species have been reported from the western equatorial Atlantic (Demerara Rise) (P. Sexton, pers. comm., 2004). [Coxall & Pearson 2006]
Distinguishing features: Parent taxon (Clavigerinella): Final chambers clavate. This taxon: Final chambers paddlewheel-like, i.e. flattened in side view, sub-triangular in edge view
NB These concise distinguishing features statements are used in the tables of daughter-taxa to act as quick summaries of the differences between e.g. species of one genus. They are being edited as the site is developed and comments on them are especially welcome.
Description
Morphology: Planispiral, involute, biumbilicate; 4 - 4½ chambers in the final whorl, increasing rapidly in size as added; early chambers rounded, final 2-3 adult chambers radially elongate, distinctly flattened in the direction of coiling into subtriangular paddles, arranged perpendicular to the direction of coiling; individual chambers are wide at the distal ends, tapering to a narrow waist at the point of attachment; peripheral outline strongly lobulate; equatorial narrow arched aperture, symmetrical or slightly asymmetrical, bordered by an imperforate lip; sutures straight or curved, short compared to the total length of chambers. [Coxall & Pearson 2006] Wall type: Uncertain because all the specimens we have examined are poorly preserved; it may have a higher pore density than is typical of the genus. [Coxall & Pearson 2006] Size: Maximum diameter of holotype 0.66 mm; minimum diameter 0.51 mm; maximum thickness (at periphery) 0.40 mm; minimum thickness (near umbilicus) 0.15 mm (Petters, 1954). Specimens reach up to 1 mm in diameter. [Coxall & Pearson 2006]
Character matrix
test outline:
Lobate
chamber arrangement:
Planispiral
edge view:
Hourglass
aperture:
Equatorial
sp chamber shape:
N/A
coiling axis:
N/A
periphery:
N/A
aperture border:
Thin lip
umb chbr shape:
Subtriangular
umbilicus:
Wide
periph margin shape:
Broadly rounded
accessory apertures:
None
spiral sutures:
N/A
umb depth:
Shallow
wall texture:
-
shell porosity:
-
umbilical or test sutures:
Weakly depressed
final-whorl chambers:
4-4.5
N.B. These characters are used for advanced search. N/A - not applicable
Biogeography and Palaeobiology
Geographic distributionApparently worldwide, most common in continental margin settings and the equatorial Pacific. Not found in open-ocean oligotrophic assemblages. It was originally described from the same sample as Pseudoglobigerinellabolivariana. Blow’s (1979) comment that this species is particularly common in the Indo-Pacific region has not been substantiated. [Coxall & Pearson 2006]
Aze et al. 2011 summary: Cosmopolitan on continental shelf settings; based on Coxall & Pearson (2006) Isotope paleobiologyClavigerinellacolombiana registers high δ18O and low δ13C compared to other co-occurring planktonic species, indicating that it lived in a cold, 12C-rich water mass (Coxall, 2000). [Coxall & Pearson 2006] Aze et al. 2011 ecogroup 4 - Open ocean sub-thermocline. Based on very light _13C and very heavy _18O. Sources cited by Aze et al. 2011 (appendix S3): Coxall et al. (2000) Phylogenetic relationsThe evolutionary relationships of C. colombiana to other clavigerinellids is difficult to establish because of its sporadic occurrence. It probably evolved from C. eocanica in the latest early Eocene (E7) by lateral widening and flattening of the chambers in the direction of coiling. [Coxall & Pearson 2006]
Most likely ancestor:Clavigerinella eocanica - at confidence level 3 (out of 5). Data source: Coxall & Pearson (2006), fig 8.1.
Biostratigraphic distribution
Geological Range: Notes: Uppermost Zone E7-E10 (poorly constrained). [Coxall & Pearson 2006] Last occurrence (top): at top of E10 zone (100% up, 41.9Ma, in Lutetian stage). Data source: Coxall & Pearson (2006), fig 8.1 First occurrence (base): in upper part of E7a subzone (80% up, 48.7Ma, in Ypresian stage). Data source: Coxall & Pearson (2006), fig 8.1
Plot of occurrence data:
Range-bar - range as quoted above, pink interval top occurs in, green interval base occurs in.
Triangles indicate an event for which a precise placement has been suggested
(NB There is no histogram as there are no occurrence records for the taxon in the Neptune database) Parent: Clavigerinella
Primary source for this page: Coxall & Pearson 2006 - Eocene Atlas, chap. 8, p. 221
References:
Blow, W. H. (1979). The Cainozoic Globigerinida: A study of the morphology, taxonomy, evolutionary relationships and stratigraphical distribution of some Globigerinida (mainly Globigerinacea). E. J. Brill, Leiden. 2: 1-1413. gs
Coxall, H. K. & Pearson, P. N. (2006). Taxonomy, biostratigraphy, and phylogeny of the Hantkeninidae (Clavigerinella, Hantkenina and Cribrohantkenina). In, Pearson, P. N., Olsson, R. K., Hemleben, C., Huber, B. T. & Berggren, W. A. (eds) Atlas of Eocene Planktonic Foraminifera. Cushman Foundation for Foraminiferal Research, Special Publication . 41(Chap 8): 213-256. gsO
Coxall, H. K. (2000). Hantkeninid planktonic foraminifera and Eocene palaeoceanographic change. In, p264 (ed.) . PhD thesis, University of Bristol (unpublished): 1-264. gs
Petters, V. (1954). Tertiary and Upper Cretaceous foraminifera from Colombia, S. A. Contributions from the Cushman Foundation for Foraminiferal Research. 5(1): 37-41. gs
Weiss, L. (1955b). Planktonic index foraminifera of northwestern Peru. Micropaleontology. 1: 301-319. gs
Clavigerinella colombiana compiled by the pforams@mikrotax project teamviewed: 7-12-2024
Eocene_Atlas_pl08-2_fig11-12.jpg
Eocene_Atlas_pl08-2_fig13.jpg
Eocene_Atlas_pl08-2_fig11-12.jpg
Eocene_Atlas_pl08-2_fig13.jpg
holotype: H. colombiana
holotype: H. colombiana
