Globorotaliabroedermanni Cushman and Bermúdez.— Toumarkine, 1975:738, pl. 2: figs. 17, 18 [middle Eocene Globorotalialehneri Zone, DSDP Site 313, Shatsky Rise, northwestern Pacific Ocean].—Toumarkine and Luterbacher, 1985:130, text-figs. 17-18 (reillustration of Toumarkine, 1975, pl. 2: figs. 17, 18); ?19 [ middle Eocene G. lehneri Zone, DSDP Site 305, Central Pacific Mountains, Central Pacific Ocean]. [Not Cushman and Bermúdez, 1949.]
Globorotalia (Acarinina) broedermanni anapetes Blow, 1979:914-915, pl. 172: figs. 1-5, 6 (holotype) [middle Eocene Zone P11, Sample RS.24, Kilwa area, Tanzania, East Africa].
Igorinaanapetes (Blow)._Pearson and others, 2004:37, pl. 2, fig. 3 [middle Eocene, Zone P11, Tanzania Drilling Project Site 2, Kilwa Masoko, Tanzania]; Berggren et al. 2006, p. 380.
Taxonomic discussion: Blow (1979, p. 914) distinguished this taxon from the ancestral broedermanni and viewed it as the terminal member of the convexa -broedermanni lineage. Aside from the specimens illustrated by Blow (1979, pl. 172) and the 9-chambered form from the G. lehneri Zone at DSDP 313 (Central Pacific) illustrated by Toumarkine (1975, pl. 2, figs. 17,18), we have not come across valid references to this form in the literature. The taxon is, however, moderately abundant in Tanzanian drill-cores that were recently taken near the type locality (Pearson and others, 2004). [Berggren et al. 2006]
Distinguishing features: Parent taxon (Pearsonites): small Morozovellids with biconvex–planoconvex tests with an overall rounded periphery This taxon: Like I. broedermanni but with more plano-convex test; more chambers (8-9) in final whorl; wider and deeper umbilicus; and tendency towards an intraumbilically restricted aperture.
NB These concise distinguishing features statements are used in the tables of daughter-taxa to act as quick summaries of the differences between e.g. species of one genus. They are being edited as the site is developed and comments on them are especially welcome.
Description
Diagnostic characters: Distinguished from broedermanni by its more pronouncedly plano-convex test, larger number of chambers (8-9) in final whorl, wider, deeper umbilicus, and tendency towards an intraumbilically restricted aperture. [Berggren et al. 2006] Morphology: Test low trochospiral, subcircular, weakly lobulate; in umbilical view 8-9 almost equidimensional, subtriangular chambers, intercameral sutures nearly straight, radial, moderately incised, umbilicus relatively broad and deep, aperture a continuous, circum-umbilical low arching slit (exposing earlier chambers in the umbilical region), extending towards (but not as far as) the peripheral margin, bordered by a thin lip; in spiral view about 18-20 chambers arranged in 2½ to 3 whorls, intercameral sutures flush with test, proximally radial, becoming retorse towards junction with peripheral margin; in edge view plano-convex; peripheral margin rounded to subacute with no evidence of peripheral concentration of muricae. [Berggren et al. 2006] Wall type:
Soldan et al 2014 f05-3d.JPG
Soldan et al 2014 f05-3e.JPG
Soldan et al 2014 f07-3d.JPG
Soldan et al 2014 f08-d.JPG
Soldan et al 2014 f08-e.JPG
Densely muricate, normal perforate, nonspinose. [Berggren et al. 2006] Size: Dimensions of holotype: maximum diameter: 0.32 mm (Blow, 1979, p. 914). [Berggren et al. 2006]
Character matrix
test outline:
Subcircular
chamber arrangement:
Trochospiral
edge view:
Planoconvex
aperture:
Umbilical-extraumbilical
sp chamber shape:
Inflated
coiling axis:
Low
periphery:
N/A
aperture border:
Thin lip
umb chbr shape:
Subtriangular
umbilicus:
Wide
periph margin shape:
Narrowly rounded
accessory apertures:
None
spiral sutures:
Weakly depressed
umb depth:
Deep
wall texture:
Coarsely muricate
shell porosity:
Finely Perforate: 1-2.5µm
umbilical or test sutures:
Weakly depressed
final-whorl chambers:
8-9
N.B. These characters are used for advanced search. N/A - not applicable
Biogeography and Palaeobiology
Geographic distributionLimited citation hinders determination of its geographic distribution; reported from East Africa and the north-west Pacific (see synonymy above); probably widely distributed in (sub)tropical areas of the world. [Berggren et al. 2006]
Aze et al. 2011 summary: Low to middle latitudes; based on Berggren et al. (2006a) Isotope paleobiologyNo data available. [Berggren et al. 2006] Aze et al. 2011 ecogroup 1 - Open ocean mixed-layer tropical/subtropical, with symbionts; based on comparison with other species of the genus. Phylogenetic relationsLineal descendant of Igorinabroedermanni and apparently the end-member of the igorinid lineage. [Berggren et al. 2006]
Most likely ancestor: Pearsonites broedermanni - at confidence level 4 (out of 5). Data source: Berggren et al. (2006), f12.1.
Biostratigraphic distribution
Geological Range: Notes: Zones E8-top of E9. We have also found in borehole TDP-13 in Tanzania (Pearson, P. N., personal observation) that anapetes has its LAD at the same level as Morozovellaaragonensis (i.e., at the Zone E9/10 boundary). [Berggren et al. 2006] Last occurrence (top): at top of E9 zone (100% up, 43.2Ma, in Lutetian stage). Data source: Berggren et al. (2006), f12.1 First occurrence (base): at base of E8 zone (0% up, 45.7Ma, in Lutetian stage). Data source: Berggren et al. (2006), f12.1
Plot of occurrence data:
Range-bar - range as quoted above, pink interval top occurs in, green interval base occurs in.
Triangles indicate an event for which a precise placement has been suggested
Histogram - Neptune occurrence data from DSDP and ODP proceedings. Pale shading <50 samples in time bin. Interpret with caution & read these notes
Taxon plotted: Pearsonites anapetes, synonyms included - Igorina anapetes;
Primary source for this page: Berggren et al. 2006 - Eocene Atlas, chap. 12, p. 380
References:
Berggren, W. A., Olsson, R. K. & Premoli Silva, I. (2006a). Taxonomy, biostratigraphy and phylogenetic affinities of Eocene Astrorotalia, Igorina, Planorotalites, and Problematica (Praemurica? lozanoi). In, Pearson, P. N., Olsson, R. K., Hemleben, C., Huber, B. T. & Berggren, W. A. (eds) Atlas of Eocene Planktonic Foraminifera. Cushman Foundation for Foraminiferal Research, Special Publication . 41(Chap 12): 377-400. gs
Bermudez, P. J. (1949). Tertiary smaller foraminifera of the Dominican Republic. Cushman Laboratory for Foraminiferal Research, Special Publication. 25: 1-322. gs
Blow, W. H. (1979). The Cainozoic Globigerinida: A study of the morphology, taxonomy, evolutionary relationships and stratigraphical distribution of some Globigerinida (mainly Globigerinacea). E. J. Brill, Leiden. 2: 1-1413. gs
Pearson, P. N., et al. (2004). Paleogene and Cretaceous sediment cores from the Kilwa and Lindi areas of coastal Tanzania: Tanzania Drilling Project Sites 1–5. Journal of African Earth Sciences. 39: 25-62. gs
Soldan, D. M., Petrizzo, M. R., Silva, I. P. & Cau, A. (2011). Phylogenetic relationships and evolutionary history of the Paleogene genus through parsimony analysis. Journal of Foraminiferal Research. 41: 260-284. gs
Soldan, D. M., Petrizzo, M. R. & Silva, I. P. (2014). Pearsonites, a new Paleogene planktonic foraminiferal genus for the broedermanni lineage. Journal of Foraminiferal Research. 44: 17-27. gs
Toumarkine, M. & Luterbacher, H. (1985). Paleocene and Eocene planktic foraminifera. In, Bolli, H. M., Saunders, J. B. & Perch-Neilsen, K. (eds) Plankton Stratigraphy. Cambridge Univ. Press, Cambridge 87-154. gs
Toumarkine, M. (1975). Middle and Late Eocene planktonic foraminifera from the northwestern Pacific Ocean: Leg 32 of the Deep Sea Drilling Project. Initial Reports of the Deep Sea Drilling Project. 32: 735-751. gs
Pearsonites anapetes compiled by the pforams@mikrotax project teamviewed: 8-2-2026