pforams@mikrotax - Quiltyella clavacella

Quiltyella clavacella

Classification: pf_cenozoic -> Globigerinidae -> Quiltyella -> Quiltyella clavacella
Sister taxa: Q. clavacella, Q. nazcaensis, Q. sp.


Citation: Quiltyella clavacella (Rögl, 1969)
Rank: species
Basionym: Hastigerinella clavacella Rögl, 1969

Taxonomic discussion:

This species has been described for the Karpatian regional stage of the Paratethys, corresponding to Zone M4. Two of the specimens illustrated by Popescu and Brotea (1989) as Protentella rohiensis (holotype nr 105.560, pl. I. figs. 1, 2 and paratype P.105.561, pl. I. figs. 4, 5) are here placed in synonymy with Q. clavacella because they show the typical radially elongated and digitate chambers in the last whorl, ending with a spherical termination. Like the globigerinellid digitate forms Q. clavacella occurs only sporadically and its range is difficult to determine. Based on the range of P. rohiensis in Popescu and Brotea (1989) and the specimens from the Nazca Plate region of the Pacific we extend the range of Q. clavacella to Zone O4. [Spezzaferri et al. 2018]

Catalog entries: Hastigerinella clavacella, Protentella rohiensis

Type images:

Distinguishing features:
Parent taxon (Quiltyella): Like Globigerinella but adult chambers radially elongate. (Oligocene - Mid Miocene)
This taxon: Like nazcaensis but chambers shorter and without inflated ends

NB These concise distinguishing features statements are used in the tables of daughter-taxa to act as quick summaries of the differences between e.g. species of one genus.
They are being edited as the site is developed and comments on them are especially welcome.


Wall type: Spinose; spines are supported by spine collars which coalesce to form ridges. Pore concentrations range from around 120 pores/50 μm2 test surface area and pore diameters range from around 1.5-2 μm.

Morphology: Test consisting of 3 whorls, initially trochospiral, tending to became irregularly coiled in the last whorl of adult specimens, 5-6 chambers strongly radially elongated, digitate and ending with a spherical termination, final chamber may be smaller than the penultimate, bulla like; sutures straight and depressed in juvenile specimens, not clearly visible in adult specimens; aperture variable in shape and position: at the base of the last chamber interiomarginal, extending around the peripheral margin, or a symmetrical arch, bordered by a distinct rim; in edge view chambers strongly elongated in shape. [Spezzaferri et al. 2018]

Size: Diameter of the holotype around 0.7-0.8 mm. [Spezzaferri et al. 2018]

Character matrix
test outline:Stellatechamber arrangement:Pseudoplanispiraledge view:Concavo-convexaperture:Extraumbilical-peripheral
sp chamber shape:Elongatecoiling axis:Very lowperiphery:N/Aaperture border:Thin lip
umb chbr shape:Elongateumbilicus:Wideperiph margin shape:Broadly roundedaccessory apertures:None
spiral sutures:Moderately depressedumb depth:Shallowwall texture:Cancellateshell porosity:Macroperforate: >2.5µm
umbilical or test sutures:Moderately depressedfinal-whorl chambers:5.0-6.0 N.B. These characters are used for advanced search. N/A - not applicable

Biogeography and Palaeobiology

Geographic distribution: This species has been found in the Molasse Basin of the Austrian Paratethys and in the east Pacific Ocean (DSDP Hole 320B). [Spezzaferri et al. 2018]

Isotope paleobiology: No data available. [Spezzaferri et al. 2018]

Phylogenetic relations: The phylogenetic relationship is uncertain because of limited stratigraphic information and sporadic occurrences of the morphotypes. Quiltyella clavacella may have evolved from G. roeglina n. sp. in mid-Oligocene Zone O4 (transitional forms of G. roeglina/Q. clavacella are in Plate 6.12, Figs. 10-11). Alternatively, the irregular coiling is suggestive of a connection with G. navazuelensis. We suggest Q. clavacella gave rise to Q. nazcaensis at the Zone O6/O7 boundary. [Spezzaferri et al. 2018]

Most likely ancestor: Globigerinella roeglina - at confidence level 1 (out of 5). Data source: Spezzaferri et al. 2018.
Likely descendants: Quiltyella nazcaensis; plot with descendants

Biostratigraphic distribution

Geological Range:
Notes: Lower Oligocene Zone O4 to the top of the lower Miocene Zone M4 (Rögl, 1969). [Spezzaferri et al. 2018]
Last occurrence (top): within M4 zone (16.38-17.54Ma, top in Burdigalian stage). Data source: Spezzaferri et al. 2018
First occurrence (base): within O4 zone (28.09-29.18Ma, base in Rupelian stage). Data source: Spezzaferri et al. 2018

Plot of occurrence data:

Primary source for this page: Spezzaferri et al. 2018 - Olig Atlas chap.6 p.207


Cicha, I., Rögl, F., Rupp, C. & Ctyroká, J. (1998). Oligocene-Miocene foraminifera of the central Paratethys. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft. 549: 1-325. gs

Popescu, G. & Brotea, D. (1989). Genus Protentella (Foraminifera) in North Transylvania Oligcene. In, Petrescu, I. (ed.) The Oligocene from the Transylvanian Basin. Cluj-Napoca 255-260. gs

Rögl, F. (1969b). Die Miozäne Foraminiferenfauna von Laa an der Thaya in der Molassezone von Niederösterreich. Mitteilungen der Geologischen Gesellschaft in Wien. 61: 63-123. gs

Spezzaferri, S., Coxall, H. K., Olsson, R. K. & Hemleben, C. (2018a). Taxonomy, biostratigraphy, and phylogeny of Oligocene Globigerina, Globigerinella, and Quiltyella n. gen. In, Wade, B. S., Olsson, R. K., Pearson, P. N., Huber, B. T. & Berggren, W. A. (eds) Atlas of Oligocene Planktonic Foraminifera. Cushman Foundation for Foraminiferal Research, Special Publication. 46(Chap 6): 179-214. gs V O


Quiltyella clavacella compiled by the pforams@mikrotax project team viewed: 7-12-2021

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