pforams@mikrotax - Woodringina hornerstownensis

Woodringina hornerstownensis

Classification: pf_cenozoic -> Guembelitrioidea -> Guembelitriidae -> Woodringina -> Woodringina hornerstownensis
Sister taxa: W. hornerstownensis, W. claytonensis, W. sp.


Citation: Woodringina hornerstownensis Olsson 1960
Rank: Species
Basionym: Woodringina hornerstownensis
Taxonomic discussion: The holotype illustration and description of Chiloguembelina taurica Morozova, 1961, closely resembles W. homerstownsis Olsson, 1960. Chiloguembelina taurica was originally described as being characterized by a "high narrow test, weakly compressed bilaterally, its height two to three times the width. Lateral outlines at first subtriangular, later almost parallel" ( Morozova, 1961:18). The test of C. taurica is formed of 10-12 spheroidal chambers, and its intercameral sutures are almost straight. Although not visible in the holotype illustrations, the aperture of C. taurica was described as semicircular and basal ( Morozova, 1961). Based on its type illustration and description, Chiloguembelina taurica Morozova, 1961, should be considered a possible junior synonym of Woodringina hornerstownensis Olsson, 1960. The holotype illustrations and descriptions of Heterohelix gradata Khalilov, 1967, and Heterohelix gradata normalis Khalilov, 1967, also closely resemble Woodringina hornerstownensis Olsson, 1961. Heterohelix gradata normalis was originally described as being characterized by an elongate "wedge-shaped [test], gradually broadening toward the obliquely trimmed-off apertural end ... chambers biserially arranged, in each offset row there are 6-8 spherical chambers. ... Aperture semilunate, much shifted from the median frontal position and shielded on one side by a moderately protruding lip" (Khalilov, 1967:173). From this description H. gradata normalis is indistinguishable from W. hornerstownensis in general test morphology. Furthermore, it appears to share the asymmetry of apertural shape and position that is diagnostic of Woodringina and related taxa. Heterohelix gradata sensu stricto was distinguished from H. gradata normalis by its last 4-10 chambers being much larger than the preceding chambers (Khalilov, 1967). It also appears to share the chamber shape, adult chamber arrangement, and apertural asymmetry of W. hornerstownensis. Given the general congruence of their original illustrations and descriptions with W. hornerstownensis, Heterohelix gradata Khalilov and H. gradata normalis Khalilov should be considered possible junior synonyms of W. hornerstownensis Olsson, 1960. The original descriptions of H. gradata and H. gradata normalis stated that the tests of these taxa are covered with large pores (Khalilov, 1967). If so, these taxa are distinguishable from W. hornerstownensis on the basis of wall structure.
Close examination of wall structure and other relevant characters (i.e., the presence or absence of initial triseriality) in type populations of H. gradata and H. gradata normalis are necessary to conclusively define the taxonomic status of these taxa relative to W. hornerstownensis. [Olsson et al. 1999]

Catalog entries: Woodringina hornerstownensis, Chiloguembelina taurica

Type images:

Distinguishing features:
Parent taxon (Woodringina): Tests contain an initial whorl of 3 chambers, later whorls of 2 chambers each. Test wall microperforate, marked by a guembelitriid surface texture (smooth walled or bearing perforate pustules). Aperture usually asymmetrically positioned and thin apertural lip infolded on one side.
This taxon: Test biserial, elongate, tapering with 6 or more chamber pairs, distinctly twisted. Wall microperforate. Aperture with a thin lip. Apertural position and height are somewhat variable.

NB These concise distinguishing features statements are used in the tables of daughter-taxa to act as quick summaries of the differences between e.g. species of one genus.
They are being edited as the site is developed and comments on them are especially welcome.


Emended description:


Wall type:


Character matrix
test outline:Triangularchamber arrangement:Biserialedge view:Equally biconvexaperture:Terminal
sp chamber shape:N/Acoiling axis:N/Aperiphery:N/Aaperture border:Thin flange
umb chbr shape:Inflatedumbilicus:N/Aperiph margin shape:Broadly roundedaccessory apertures:None
spiral sutures:N/Aumb depth:N/Awall texture:Finely pustuloseshell porosity:Microperforate: <1┬Ám
umbilical or test sutures:Moderately depressedfinal-whorl chambers:2.0-2.0 N.B. These characters are used for advanced search. N/A - not applicable

Biogeography and Palaeobiology

Geographic distribution: Widespread in high and low latitudes (Figure 34). [Olsson et al. 1999]

Isotope paleobiology: Biogeographic and stable isotopic data suggest an open-ocean, warm shallow-water habitat for W. hornerstownensis, similar to that of W. claytonensis (D'Hondt and Keller, 1991; Liu and Olsson, 1992; D'Hondt and Zachos, 1993). [Olsson et al. 1999]

Phylogenetic relations: This species is generally considered to have descended from Guembelitria cretacea via W. claytonensis (Olsson, 1970, 1982; Smit, 1977, 1982; D'Hondt, 1991; Li and Radford, 1991; Olsson et al., 1992; Liu and Olsson, 1992). [Olsson et al. 1999]

Most likely ancestor: Woodringina claytonensis - at confidence level 0 (out of 5). Data source: .
Likely descendants: Chiloguembelina morsei; plot with descendants

Biostratigraphic distribution

Geological Range:
Notes: Zone P a to Zone P3b. [Olsson et al. 1999]
Last occurrence (top): within P3b subzone (60.73-61.33Ma, top in Selandian stage). Data source: Olsson et al. 1999
First occurrence (base): within Pa zone (65.72-66.00Ma, base in Danian stage). Data source: Olsson et al. 1999

Plot of occurrence data:

Primary source for this page: Olsson et al. 1999 - Atlas of Paleocene Planktonic Foraminifera, p. 87


D'Hondt, S. & Keller, G. (1991). Some patterns of planktic foraminiferal assemblage turnover at the Cretaceous-Tertiary boundary. Marine Micropaleontology. 17: 77-118. gs

D'Hondt, S. & Zachos, J. C. (1993). On Stable Isotopic Variation and Earliest Paleocene Planktonic Foraminifera. Paleoceanography. 8: 527-547. gs

Liu, C. & Olsson, R. K. (1992). Evolutionary radiation of microperforate planktonic foraminifera following the K/T mass extinction event. Journal of Foraminiferal Research. 22: 328-346. gs

Olsson, R. K. (1960). Foraminifera of late Cretaceous and earliest Tertiary age in the New Jersey Coastal Plain. Journal of Paleontology. 34(1): 1-58. gs

Olsson, R. K., Hemleben, C., Berggren, W. A. & Huber, B. T. (1999). Atlas of Paleocene Planktonic Foraminifera. Smithsonian Institution Press, Washington, DC. 1-252. gs


Woodringina hornerstownensis compiled by the pforams@mikrotax project team viewed: 7-10-2022

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