1899 Hexadoras borealis - Cleve p. 15, 30; pl. 2, fig. 4
1900 Rhizoplegma boreale - Jørgensen p. 61-62
1905 Rhizoplegma boreale - Jørgensen p. 118, pl. 9, fig. 38; pl. 10, fig. 38
1908 Rhizoplegma boreale var antarctica - Popofsky p. 216-217, pl. 24, fig. 1
1909 Rhizoplegma boreale - Schröder fig. 23
1937 Rhizoplegma boreale - Wailes fig. 20
1952 Rhizoplegma boreale antarctica - Dogiel and Reschetnjak p. 8
1967 Rhizoplegma (?) boreale - Petrushevskaya p. 12-14, fig. 8
1971 Rhizoplegma boreale - Ling et al. p. 710, fig. 3; pl. 1, figs. 2-3
1976 Rhizoplegma boreale - Bjørklund pl. 3, figs. 10-16; pl. 4, figs. 1-3
1983 Rhizoplegma boreale - Blueford tabs. 1-2, not figured
1985 Rhizoplegma boreale - Bjørklund fig. 1
1986 Rhizoplegma boreale - Swanberg and Bjørklund figs. 3-5
1987 Rhizoplegma boreale - Swanberg and Bjørklund fig. 4i; tabs. 1-3
1989 Rhizoplegma boreale - Kruglikova p. 468
1992 Rhizoplegma boreale - Abelmann pl. 1, fig. 13; tab. 1
1992 Rhizoplegma boreale - Swanberg and Eide fig. 4
1995 Rhizoplegma boreale - Itaki and Takahashi fig. 10b; tab. 1
1995 Rhizoplegma boreale - Samtleben et al. not figured
1995 Rhizoplegma boreale - Schröder-Ritzrau pl. 2, fig. 7
1997 Rhizoplegma boreale - Abelmann and Gowing tab. 3
1997 Rhizoplegma boreale - Nishimura et al. pl. 1, figs. 1-3
1997 Rhizoplegma boreale - Takahashi tab. 1
1998 Rhizoplegma boreale - Bjørklund et al. pl. 1, fig. 8
1998 Rhizoplegma boreale - Dolven pl. 9, figs. 2-3
From Suzuki Paleotax db:
1899 Hexadoras borealis n. sp. Cleve p. 30, pl. 2, fig. 4a, 4b, 4c
1899 Rhizoplegma boreale (Cleve) Jörgensen p. 61-62
1905 Rhizoplegma boreale (Cleve) Jörgensen p. 118, pl. 9, fig. 38, pl. 10, fig. 38e, 38f
1937 Rhizoplegma boreale (Cleve) Wailes p. 11, fig. 20
1952 Rhizoplegma borealis atalntica (Cleve) [sic] Dogiel' & Reshetnyak p. 8
1967 Rhizoplegma (?) boreale (Cleve) Petrushevskaya p. 16-17, text-fig. 8.1, 8.2
1969 Rhizoplegma boreale (Cleve) Petrushevskaya pl. 3, fig. 7
1971 Rhizoplegma boreale (Cleve) Ling et al. p. 710, pl. 1, figs. 2, 3
1974 Rhizoplegma boreale (Cleve) Ling p. 9, fig. 7
1975 Rhizoplegma boreale (Cleve) Aarseth et al. fig. 14c
1976 Rhizoplegma boreale (Cleve) Bjørklund pl. 3, figs. 10-16, pl. 4, figs. 1-3
1977 Rhizoplegma boreale (Cleve) Kling p. 217, pl. 1, fig. 3
1984 Rhizoplegma boreale (Cleve) Nishimura & Yamauchi p. 32, pl. 15, fig. 7
1986 Rhizophlegma boreale (Cleve) [sic] Bjørklund & Goll pl. 1, figs. 3, 4
1990 Hexacladus sp. Fujioka pl. 39, fig. 8
2013 Cleveiplegma borealis (Cleve) Dumitrica p. 25, pl. 1, figs. 1-9
2014 Cleveiplegma boreale (Cleve) Matsuzaki et al. pl. 1, fig. 11
2015 Cleveiplegma borealis (Cleve) Matsuzaki et al. p. 16, figs. 3.1, 3.2
2017 Rhizoplegma boreale (Cleve) Chen et al. p. 122-123, pl. 27, figs. 1-14
Catalog entries: Hexadoras borealis
Original description: Primordial shell: irregularly spherical, 30-40µm. in diameter, with irregular, rounded or polygonal pores, 2 to 3 on the radius, and thin bars. Spines six, exceptionally more, strong, with triangular apophyses in the middle. Pl. II, fig. 4 a. 0uter shell: a rounded or octahedric, more or less intricate net-work of anastomosing, silicious threads, issuing from the proximal edges of the spines. Spines usually six (rarely as in Rhizoplegma 8 to 10) strong, three-sided slightly spirally twisted, with elegantly aculeate, winged edges.
Diam. 120-160µm. Spines 100µm.
Surface.
Jørgensen 1905 - Hexadoras borealis Easily recognized by the characteristic interwoven pyramids around the main spines. Haeckel depicts similar pyramids for Rhizoplegma lychnosphra. The inside shell is irregular, not quite round; sometimes resembling a cube in shape, sometimes it is more like an octahedron. The pores are uneven, irregular, polygonal, averaging about 10µm. The intermediate walls between the pores are not broad, differently developed, but never having the distinct broader corners which correspond to the rounded lumen of the pore. A few small, short, needle-shaped byspines are found here and there on the beams, but not regularly in the corners. The main spines are long and strong, often 8 in number (according to Haeckel's system answering to an inner cube), although also often only 6 (answering to an octahedron) or 7. A larger number may sometimes be found. The main spines have 3 strongly developed edges which narrow off toowards both ends. On these edges there are transverse branches diverging at right angels and these, together with the corresponding ones on the other edges of the same spine, form a very variable number of verticils of threes. The spongy, loose and very irregularly constructed network, which forms the outer shell, arises from the lowest 2-3 verticils on the main spines, but has the appearance of being lengthened out a good way up along the radial spines, on account of the thin connecting beams, which unite the different verticils parallel to the direction of the spine. Also on the outside of the outer spongy shell (network), there are some few scattered short and fine, needle-shaped byspines. The ocean forms seem generally to have 6 radial spines. There seems to be a slight difference between the regular forms which have 6 radial spines, and those which are more frequently found in the coast water and which have about 8. The former seem to have a smaller inner shell with distinct byspines. I have, however, not discovered any definite difference so as to make it necessary to divide them into two or more species. Young forms are rather unrecognizable, as both the interwoven pyramids and the spongy outer shell are absent. The construction of this species resembles greatly that of Rhizoplegma radicatum HCK and Rhizoplegma lychnosphæra HCK. The inner, fine, transverse branches on the radial spines in the space between the outer, spongy shell and the inner shell are, however, wanting. These transverse branches are by Haeckel the characteristic of the subgenus Rhizoplegmidium. According to Haeckel's system, it might perhaps be a question as to whether or not our species should be classed as belonging to the genus Lychnosphæra, because of the byspines on the inner shell. As these, however, are small and few in number, and perhaps not even always present, this would not be recommendable, and still more so as such a distinction between the genera seems to be unnatural. Haeckel's genus Hexadoras has radial spines without side branches, for which reason I still mean that the above species finds a more natural place among the closely allied forms of the genus Rhizoplegma, notwithstanding that there undoubtedly often occur forms with 6 main spines. Belongs to the most common radiolaria with us and is not rare in deep water samples, although never numerous. Distribution: On the west coast of Norway rather frequent, although rare in fully developed condition. Known from the sea west and south of Spitzbergen (Cleve, 1899) and from a few places in the Norwegian Ocean. Petrushevskaya 1967 - Hexadoras borealis Skeleton consists of small inner sphere, radial needles and fine-lattice skeletal mass. Inner sphere of irregular shape, with irregularly disposed pores (5-7 on half equator). From it radiate 8-10 stout trihedral radial needles. Processes arise at right angles from each facet of needle; from these apophyses develop, uniting with each other and forming irregular lattices of basic skeletal mass. Between inner sphere and this lattice is small space. Radiating needles have processes in form of small barbs along entire length, but lattice is formed at first from beyond lattice. Lattice not of uniform thickness, being more developed around radial needles, where its layer is thicker, and becoming thinner in spaces between needles. Dimensions: diameter of inner sphere 30-40µm, space between internal sphere and lattice 15-20µm, diameter of chamber formed 55-70µm, thickness of layer of lattice mass 30-50µm, diameter of lattice space 10-15µm, width of radial needles 8-10µm.
Published descriptions
Geological Range:
Last occurrence (top): Extant. Data source: Lazarus et al. 2015 - "R age group"
First occurrence (base): within Quaternary Period (0.00-2.59Ma, base in Gelasian stage). Data source: Lazarus et al. 2015 - "R age group"
Plot of occurrence data:
Bjørklund, K. R. (1976). Radiolaria from the Norwegian Sea, Leg 38 of the Deep Sea Drilling Project. Initial Reports of the Deep Sea Drilling Project. 38: 1101-1168. gs Bjørklund, K. R. (1985). Upper Weichselian-Holocene radiolarian stratigraphy in the Skagerrak (NE North Sea). Norsk Geologisk Tidsskrift. 65: 103-106. gs Blueford, R (1983). Distribution of Quaternary radiolarian in the Narvarin Basin geologic province, Bering Sea. Deep-Sea Research. 30(7A): 763-781. gs Dogel, V. A. & Reshetnyak, V. V. (1952). Materialy po radiolyariyam severo-zapadnoy chasti tikhogo okeana. [Material on radiolarians of the northwestern part of the Pacific Ocean]. Issledovanya Dalnevostochnykh Morei SSSR. 3: 5-36. gs Jørgensen, E (1900). Protophyten und Protozoen im Plankton aus der norwegischen Westkuste. Bergens Museums Aarbog. 1899(6): 51-95. gs O Jørgensen, E (1905). The protist plankton and diatoms in bottom samples: Radiolaria. In, Nordgaard (ed.) Hydrographical and biological investigation in Norwegian Fjord. 49-151. gs O Ling, H-y., Stadum, C. J. & Welch, M. L. (1971). Polycystine Radiolaria from Bering Sea surface sediments. In, Farinacci, A. (ed.) Proceedings of the II Planktonic Conference, Roma 1970. Tecnoscienza, Roma 705-729. gs Petrushevskaya, M. G. (1967). Radiolayarii otryadov Spumellaria i Nassellaria Antarkicheskoi oblasti (po materialam Sovetskoi Antarkitcheskikh Ekspeditsii). [Antarctic Spumelline and Nasselline radiolarians (from materials from the Soviet Antarctic Expedition)]. Issledovanie Faunyi Morey (Akademii Nauk SSSR) Leningrad. 4(12): 5-186. gs Popofsky, A (1908). Die Radiolarien der Antarktis (mit Ausnahme der Tripyleen). Deutsche Sudpolar-Expedition 1901-1903. 10(3): 183-305. gs O Schröder, O. (1909a). Die nordischen Spumellarian. In, Brandt, K. & Apstein, C. (eds) Nordisches Plankton. Lipsius und Tischer, Kiel and Leipzig, Germany (17): 1-66. gs O Swanberg, N. R. & Bjørklund, K. R. (1986). The radiolarian fauna of western Norwegian fjords; patterns of abundance in the plankton. Marine Micropaleontology. 11: 231-241. gs Swanberg, N. R. & Bjørklund, K. R. (1987a). Radiolaria in the plankton of some fjords in western and northern Norway: the distribution of species. Sarsia. 72(3-4): 231-244. gs Wailes, G. H. (1937). 1. Protozoa. 1a. Lobosa, 1b. Reticulosa, 1c. Heliozoa, 1d. Radiolaria. University of Toronto Press for the Biological Board of Canada, Toronto. 9-14. gs Missing or ambiguous references: Cleve 1899; References:
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Rhizoplegma boreale compiled by the radiolaria@mikrotax project team viewed: 17-6-2025
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