pforams@mikrotax - Favusella washitensis pforams@mikrotax - Favusella washitensis

Favusella washitensis

Classification: pf_mesozoic -> Favusellidae -> Favusella -> Favusella washitensis
Sister taxa: F. hoterivica, F. washitensis, F. sp.


Citation: Favusella washitensis (Carsey, 1926)
taxonomic rank: Species
Basionym: Globigerina washitensis Carsey, 1926
Synonyms: Synonyms list form Koutsoukos et al 1989
Taxonomic discussion: Favusella washitensis is regarded as a species comprising several morphotypes of common origin. The different morphotypes represent different polymorphic stages in the life cycle and ecophenotypes of a single taxon.
Specimens attributed to Favusella are distinguised from Hedbergella by having a distinctly trochoid chamber shape, thicker and distinctly ornamented wall, characteristic pore distribution pattern, and umbilical position of the aperture. [copied from Chronos database]

Type images:

Original description: Shell calcareous, usually white; surface very coarsely and regularly pitted; chambers well inflated, rapidly enlarging, arranged subspirally, dorsal side shows two or more whorls; ventral side shows only one whorl which consists of the last three or four chambers rather loosely joined about a depressed center; aperture at the inner margin of the final chamber. 

Entries in the Catalog of original descriptions: Globigerina washitensis, Favusella confusa, Favusella hedbergellaeformis, Hedbergella hiltermanni, Favusella nitida, Favusella orbiculata, Favusella papagayosensis, Favusella pessagnoi, Favusella planata, Favusella quadrata, Favusella scitula, Ascoliella scotiensis, Favusella stiftia, Favusella voloshinae, Favusella wenoensis

Emended description:

Emended : Test a low to relatively high trochospiral coil, equatorial periphery lobate, peripheral edge rounded. Chambers spherical to subspherical; 1.5 to 3.5 whorls each with 3 to 6 chambers, usually 4 to 5 in last whorl. Last chamber in mature individuals is usually smaller (embryonic), bare or with smooth ornamentation, extending over the umbilicus and partially covering it. Primary aperture usually arcuate, low to moderate, interiomarginal, umbilical to umbilical-extraumbilical in position, bordered by wide or narrow lip. Umbilicus shallow, of variable width. Sutures radial to slightly curved, depressed. Test surface covered by variable sculptural ornamentation, progressively developing from fine rounded tubercles and well-defined rounded or elongated tubercles (some forming elongated quadrangles = ridges), to reticulate system of fine to coarse ridges forming irregular polygonal cells (honeycomb pattern) in ephebic (adult) stage. Numerous minute pores located nearly exclusively within polygonal cells and between tubercles; pores scattered on edges of cell-ridges and tubercles. Size (maximum diameter) <0.1-0.25mm in the neanic stage (1 to 2 whorls), 0.25-0.35mm in ephebic forms (2.5 to 3 whorls) and 0.35->0.5mm in gerontic specimens (3 to 3.5 whorls). [Koutsoukos et al 1989; copied from Chronos database]

Character matrix
test outline:Lobatechamber arrangement:Trochospiraledge view:Spiroconvexaperture:Umbilical
sp chamber shape:Inflatedcoiling axis:Moderate-highperiphery:N/Aaperture border:Thin lip
umb chbr shape:Inflatedumbilicus:Narrowperiph margin shape:Broadly roundedaccessory apertures:None
spiral sutures:Moderately depressedumb depth:Shallowwall texture:-shell porosity:Microperforate: <1µm
umbilical or test sutures:Moderately depressedfinal-whorl chambers:4-6 N.B. These characters are used for advanced search. N/A - not applicable

Geological Range:
Last occurrence (top): in lower part of Rotalipora cushmani zone (40% up, 96.8Ma, in Cenomanian stage). Data source: Carter & Hart, 1977
First occurrence (base): within P. rohri zone (113.22-116.60Ma, base in Aptian stage). Data source: copied from Chronos database, but zonal name updated from T. bejaouaensis to T. rohri

Plot of range and occurrence data:


Carsey, D. O. (1926). Foraminifera of the Cretaceous of central Texas. University of Texas Bulletin. 2612: 1-56. gs

Koutsoukos, E. A. M., Leary, P. N. & Hart, M. B. (1989). Favusella Michael (1972): Evidence of ecophenotypic adaptation of a planktonic foraminifer to shallow-water carbonate environments during the mid-Cretaceous. Journal of Foraminiferal Research. 19(4): 324-336. gs

Longoria, J. F. & Gamper, M. A. (1977). Albian planktoinc foraminifera from the Sabinas Basin of northern Mexico. Journal of Foraminiferal Research. 7(3): 190-215. gs

Michael, F. Y. (1972). Planktonic foraminifera from the Comanchean Series (Cretaceous) of Texas. Journal of Foraminiferal Research. 2(4): 200-220. gs

Pessagno, E. A. (1967). Upper Cretaceous planktonic foraminifera from the western Gulf Coastal Plain. Palaeontographica Americana. 5: 245-445. gs O

Postuma, J. A. (1971). Manual of planktonic foraminifera. Elsevier for Shell Group, The Hague. 1-406. gs

Missing or ambiguous references: Lutze & Pflaumann 1978;


Favusella washitensis compiled by the pforams@mikrotax project team viewed: 18-5-2024

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Comments (2)


Hello, thanks for this great database. Biostrati range update ? : Any reason to not follow e.g. Carter & Hart 1977, Kennedy 1969 who report the presence of F. washitensis in England during the Middle Cenomanian (just below the Orbirhnchia mantelliana Band which is at the upper limit of the Middle Cenomanian Turrilites costatus Zone)...

And what is the ref. of Koutsoukos 2005 mentioned above for emendation ? (Koutsoukos 1989?) ?

Carter & Hart 1977 aspects of mid-cretaceous stratigraphical micropal.

Kennedy 1969 The correlation of the Lower Chalk of South-East England

Koutsoukos 1989 Favusella Michael (1972) Evidence of ecophenotypic adaptation...

Best regards

Jeremy Young


Thank you for these comments, you are quite right the correct reference is Koutsoukos et al. 1989, and I have now corrected that. The age range data is not really something I can answer, I will ask Brian Huber to comment on this.